A new species of the genus Humboldtiana (Gastropoda, Stylommatophora, Xanthonychidae) from Sierra de Penjamo, Guanajuato, Mexico Author Mejía, Omar 0000-0002-7021-8377 Laboratorio de Variación Biológica y Evolución, departamento de Zoología, Escuela Nacional de Ciencias Biológicas, Instituto Politécnico Nacional, Mexico, Mexico Author López, Benjamín Laboratorio de Variación Biológica y Evolución, departamento de Zoología, Escuela Nacional de Ciencias Biológicas, Instituto Politécnico Nacional, Ciudad de México, Mexico text Biodiversity Data Journal 2024 2024-10-11 12 e 132797 e 132797 journal article 10.3897/BDJ.12.e132797 1AAAED54-97D1-4039-8C81-5C472EF47D58

Table 1. Species included in the Humboldtiana buffoniana species group used for comparison with the new species.

Character	H. dugesi holotype	H. salviahispanica	H. pinicola	H. potosiana	H. queretaroana
Shell size	small 27 mm height	small 22 mm height	large 38 mm height	small 24 mm height	large 37 mm height
Number of whorls	4.2	4.1-4.2	4.4-4.8	4.2	5
bands' colour pattern	dark brown visible at the interior	chestnut to dark brown visible at the interior	black visible at the interior	dark brown with a reddish tinge	absent
Shell sculpture	growth lines and granules	growth lines and granules	growth lines and granules	growth lines and granules	growth lines and granules
embryonic shell whorls	1.8	1.5-1.6	1.6-1.7	1.5	1.5
embryonic shell sculpture	1.3 smooth, then isolated granules and growth lines	1.0 smooth then almost imperceptible growth lines	smooth	smooth	smooth
Penis	globose 6.9 mm	short and stocky 7.8 mm	short and stocky 9. 6 mm	globose	NA
verge	stout covering half penis cavity	large and broadened composed of two lobes	long and extending to opening vagina with three or four lobes	NA	NA
epiphallus	long and cylindrical 11 mm	long and cylindrical 16 mm	short and stout 7 mm	short and stout	NA
flagellum	long 73 mm	short 35 mm	short 36 mm	short	NA
flagellum / penis + epiphallus	4.07 times	1.68 times	1.7-2.6 times	maybe 1 times	NA
dart size length	2 mm	1.5 mm	four darts of unequal size	NA	NA
glands height	3.2 mm	2 mm	NA	NA	NA
spermathecal duct length	52 mm	40 mm	36 mm	NA	NA
spermathecal caecum length	5.4 mm	5 mm	5.4 mm	caecum absent	NA
spermathecal length	4.7 mm	7 mm	25 mm	NA	NA
Note: Humboldtiana queretaroana is only known from the original description (Dall 1897). In H. potosiana, the size of the organs is not available (NA).