A new species of the genus Boulenophrys from South China (Anura, Megophryidae) Author Wang, Jian 0000-0003-4249-7767 Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China wangj-1994@outlook.com Author Lin, Shi-Shi 0000-0001-7709-7005 Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China Author Gan, Jun-Shen Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China Author Chen, Hong-Hui 0000-0001-5026-2920 Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China org4406chenhh@foxmail.com Author Yu, Ling-Min Guangdong Lianshan Bijiashan Provincial Nature Reserve, Qingyuan 513200, China Author Pan, Zhong Guangzhou Shengheng Eco-forestry Company, Guangzhou 510663, China Author Xiao, Jia-Jie Guangdong Wildlife Monitoring Rescue and Conservation Center, Guangzhou 510520, China Author Zeng, Zhao-Chi 0000-0003-4054-8399 Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China zhaochizeng@outlook.com text Zootaxa 2024 2024-10-01 5514 5 451 468 http://dx.doi.org/10.11646/zootaxa.5514.5.3 journal article 304414 10.11646/zootaxa.5514.5.3 07e74be0-7959-4147-902a-793b53a5d220 1175-5326 13914787 36CA84B3-FDA3-4F68-92DA-9294C2E92839

TABLE 2. Localities, voucher information, and GenBank accession numbers for Boulenophrys and Xenophrys (outgroups) samples used in this study.

ID	Species	Voucher ID	Locality	16S rRNA	CO1
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1	B. pepe sp. nov.	GEP a117	China, Guangdong, Lianshan	PQ131150	/
2	B. pepe sp. nov.	GEP a207	China, Guangdong, Lianshan	PQ131151	PQ130479
3	B. pepe sp. nov.	SYS a009350	China, Guangdong, Lianshan	PQ131152	PQ130480
4	B. pepe sp. nov.	GEP a209	China, Guangdong, Lianshan	PQ131153	PQ130481
5	B. acuta	SYS a002266	China, Guangdong, Fengkai	KJ579119	MH406122
6	B. acuta	SYS a002276	China, Guangdong, Fengkai	KJ579124	MH406126
7	B. angka	KIZ 040591	Thailand, Chiang Mai, Doi Inthanon	MN508052	/
8	B. angka	KIZ 040595	Thailand, Chiang Mai, Doi Inthanon	MN508051	/
9	B. anlongensis	CIB AL20190531018	China, Guizhou, Anlong	MT823184	MT823261
10	B. anlongensis	CIB AL20190811014	China, Guizhou, Anlong	MT823187	MT823264
11	B. baishanzuensis	CIB QY20200719002	China, Zhejiang, Qingyuan	MW001151	MT998292
12	B. baishanzuensis	CIB QY20200726002	China, Zhejiang, Qingyuan	MW001156	MT998297
13	B. baolongensis	KIZ 019216	China, Chongqing, Wushan	KX811813	KX812093
14	B. baolongensis	KIZ YPX47798	China, Chongqing, Wushan	KX811812	/
15	B. binchuanensis	SYS a006821	China, Sichuan, Yanbian	OQ180970	OQ180858
16	B. binchuanensis	KIZ 019441	China, Yunnan, Mt Jizu	KX811849	KX812112
17	B. binlingensis	FMNH 232874	China, Sichuan, Hongya	KY022205	/
18	B. binlingensis	KIZ 025807	China, Sichuan, Mt Wawu	KX811852	KX812115
19	B. boettgeri	KIZ YPXJK033	China, Fujian, Mt Wuyi	KX811814	KX812104
20	B. boettgeri	SYS a004149	China, Fujian, Mt Wuyi	MF667878	MH406247
21	B. boettgeri	SYS a002702	China, Anhui, Mt Huangshan	MF667882	MH406160
22	B. brachykolos	SYS a002441	China, Guangdong, Shenzhen	MH406691	MH406139
23	B. brachykolos	ROM 16634	China, Hong Kong	KX811897	KX812150
24	B. brachykolos	SYS a002258	China, Hong Kong	KJ560403	MH406120
25	B. caobangensis	IEBR 4385	Vietnam, Cao Bang, Nguyen Binh	LC483945	/
26	B. caobangensis	IEBR 4386	Vietnam, Cao Bang, Nguyen Binh	LC483946	/
27	B. caudoprocta	SYS a004293	China, Hunan, Sangzhi	MH406796	MH406258
28	B. caudoprocta	SYS a004308	China, Hunan, Sangzhi	MH406799	MH406261
29	B. cheni	SYS a002142	China, Hunan, Yanling	KJ560398	MH406098
30	B. cheni	SYS a004050	China, Jiangxi, Mt Jinggang	MF667873	MH406241
31	B. chishuiensis	SYS a004953	China, Guizhou, Chishui	MH406867	MH406329
32	B. chishuiensis	KIZ 025788	China, Sichuan, Mt Emei	KX811893	KX812147
33	B. chishuiensis	SYS a002164	China, Sichuan, Mt Laojun	MH406660	MH406100
34	B. congjiangensis	SYS a005134	China, Guangxi, Mt Jiuwan	MH406880	MH406342
35	B. congjiangensis	GZNU 20200706003	China, Guizhou, Congjiang	MW959773	MW959761
36	B. daiyunensis	SYS a001733	China, Fujian, Dehua	MH406643	MH406079
37	B. daiyunensis	SYS a007711	China, Fujian, Tong’an	MW367054	MW365504
38	B. daoji	SYS a004089	China, Zhejiang, Fenghua	MH406783	MH406243
39	B. daoji	SYS a006212	China, Zhejiang, Mt Tiantai	MW367047	MW365497
40	B. daweimontis	KIZ 048997	China, Yunnan, Mt Dawei	KX811867	KX812125
41	B. daweimontis	KIZ 016066	China, Yunnan, Xichou	KX811866	KX812124
42	B. daweimontis	ROM 28272	Vietnam, Lao Cai, Sapa	KX811865	/
43	B. dongguanensis	SYS a001974	China, Guangdong, Dongguan	MH406648	MH406084
44	B. dongguanensis	SYS a002007	China, Guangdong, Dongguan	MH406654	MH406090
45	B. elongata	GEP a150	China, Guangdong, Mt Lianhua	OR601592	OR597098
46	B. elongata	GEP a151	China, Guangdong, Mt Lianhua	OR601593	OR597099
47	B. fanjingmontis	KIZ 03079	China, Chongqing, Youyang	KX811859	/
48	B. fanjingmontis	KIZ 021895	China, Guizhou, Mt Fanjing	KX811860	/
49	B. fanjingmontis	SYS a004350	China, Guizhou, Mt Fanjing	MH406808	MH406270
50	B. fansipanensis	AMS R186115	Vietnam, Lao Cai, Sapa	MH514887	MW086548
51	B. fansipanensis	VNMN 2018.01	Vietnam, Lao Cai, Sapa	MH514886	MW086544
52	B. fengshunensis	SYS a004724	China, Guangdong, Fengshun	MH406848	MH406310
53	B. fengshunensis	SYS a004727	China, Guangdong, Fengshun	MH406851	MH406313
54	B. frigida	AMS R186131	Vietnam, Lao Cai, Bat Xat	MT364279	MW086550
55	B. frigida	VNMN 010948	Vietnam, Lao Cai, Bat Xat	MT364275	MW086546
56	B. hoanglienensis	VNMN 07034	Vietnam, Lao Cai, Sapa	MH514890	MW086547
57	B. hoanglienensis	VNMN 2018.02	Vietnam, Lao Cai, Sapa	MH514889	MW086551
58	B. hungtai	SYS a008576	China, Guangdong, Jiedong	OL635597	OL634864
59	B. hungtai	SYS a007577	China, Guangdong, Jiexi	OL635594	OL634861
60	B. hengshanensis	CSUFT HS210610	China, Hunan, Mt Hengshan	ON209293	/
61	B. hengshanensis	CSUFT HS210612	China, Hunan, Mt Hengshan	ON209291	/
62	B. insularis	SYS a002168	China, Guangdong, Nan’ao	MH406662	MH406102
63	B. insularis	SYS a002171	China, Guangdong, Nan’ao	MH406665	MH406105
64	B. jiangi	SYS a008382	China, Chongqing, Mt Jinfo	OQ180981	OQ180869
65	B. jiangi	KIZ 03076	China, Chongqing, Youyang	KX811911	KX812148
66	B. jiangi	CIB KKS20180722006	China, Guizhou, Kuankuoshui	MN107743	MN107748
67	B. jiangi	CIB FJS20150720004	China, Guizhou, Mt Fanjing	MN107747	MN107752
68	B. jingdongensis	MNHN 1999.5688	Vietnam, Lao Cai, Sapa	KR828082	/
69	B. jingdongensis	SYS a005160	China, Guangxi, Mt Cenwanglaoshan	MH406883	MH406345
70	B. jingdongensis	KIZ YN0705230	China, Yunnan, Mt Wuliang	KX811873	KX812134
71	B. jingdongensis	SYS a003928	China, Yunnan, Mt Wuliang	MH406773	MH406232
72	B. jinggangensis	SYS a005527	China, Hunan, Mt Dawei	OQ180963	OQ180851
73	B. jinggangensis	SYS a004824	China, Hunan, Mt Sifang	MH406857	MH406319
74	B. jinggangensis	SYS a004028	China, Jiangxi, Mt Jinggang	MH406780	MH406239
75	B. jinggangensis	SYS a003165	China, Jiangxi, Mt Jiuling	MH406740	MH406193
76	B. jiulianensis	KIZ 018496	China, Guangdong, Mt Nankun	KX811828	KX812098
77	B. jiulianensis	SYS a004011	China, Guangdong, Mt Nankun	MH406778	MH406237
78	B. jiulianensis	SYS a004218	China, Jiangxi, Mt Jiulian	MH406790	MH406252
79	B. kuatunensis	SYS a001590	China, Fujian, Mt Wuyi	KJ560377	/
80	B. kuatunensis	SYS a003449	China, Jiangxi, Mt Wuyi	MF667881	MH406206
81	B. kuatunensis	SYS a006906	China, Jiangxi, Mt Yangjifeng	OQ180971	OQ180859
82	B. leishanensis	KIZ 049172	China, Guizhou, Mt Leigong	KX811825	KX812102
83	B. leishanensis	SYS a002214	China, Guizhou, Mt Leigong	MH406674	MH406114
84	B. liboensis	GNUG 20150813001	China, Guizhou, Libo	MF285253	/
85	B. liboensis	GNUG 20160408009	China, Guizhou, Libo	MF285260	/
86	B. lini	SYS a004408	China, Hunan, Mt Bamian	MH406818	MH406280
87	B. lini	KIZ 07053	China, Jiangxi, Mt Jinggang	KX811842	KX812110
88	B. lini	SYS a002382	China, Jiangxi, Mt Jinggang	KJ560414	MH406136
89	B. lishuiensis	SYS a008445	China, Zhejiang, Liandu	OQ180984	OQ180872
90	B. lishuiensis	CIB 106670	China, Zhejiang, Liandu	KY021418	/
91	B. lushuiensis	CIB YN201909289	China, Yunnan, Lushui	MW001226	MW000913
92	B. lushuiensis	SYS a003823	China, Yunnan, Tengchong	MH406763	MH406222
93	B. lushuiensis	KIZ YPX33610	China, Yunnan, Yingjiang	KX811847	KX812142
94	B. minor	KIZ YPX37545	China, Sichuan, Dujiangyan	KX811895	KX812144
95	B. minor	SYS a003212	China, Sichuan, Dujiangyan	MF667865	MH406197
96	B. mirabilis	SYS a002192	China, Guangxi, Lingui	MH406669	MH406109
97	B. mirabilis	SYS a002289	China, Guangxi, Lingui	MH406681	MH406127
98	B. mufumontana	SYS a006419	China, Hunan, Mt Mufu	MK524107	MK524138
99	B. mufumontana	SYS a006971	China, Jiangxi, Mt Jiuling	OQ180972	OQ180860
100	B. mufumontana	SYS a005012	China, Jiangxi, Yifeng	OQ180959	OQ180847
101	B. nankunensis	SYS a004501	China, Guangdong, Mt Nankun	MH406822	MH406284
102	B. nankunensis	SYS a004503	China, Guangdong, Mt Nankun	MH406824	MH406286
103	B. nanlingensis	SYS a001962	China, Guangdong, Ruyuan	MH406645	MH406081
104	B. nanlingensis	SYS a002357	China, Jiangxi, Mt Qiyun	MH406687	MH406133
105	B. obesa	SYS a001956	China, Guangdong, Fengkai	KJ579117	/
106	B. obesa	SYS a002275	China, Guangdong, Fengkai	KJ579123	MH406125
107	B. ombrophila	CIB WY18082308	China, Fujian, Mt Wuyi	MW001159	MT998300
108	B. ombrophila	SYS a001976	China, Guangdong, Renhua	MH406650	MH406086
109	B. ombrophila	SYS a004616	China, Jiangxi, Mt Jiulian	MH406840	MH406302
110	B. omeimontis	SYS a004939	China, Guizhou, Chishui	MH406866	MH406328
111	B. omeimontis	SYS a004916	China, Sichuan, Hejiang	MH406864	MH406326
112	B. omeimontis	KIZ 025765	China, Sichuan, Mt Emei	KX811884	KX812136
113	B. omeimontis	SYS a002741	China, Sichuan, Mt Laojun	MH406710	MH406162
114	B. palpebralespinosa	FMNH 258098	Laos, Phongsaly, Phou Dendin	KY022209	/
115	B. palpebralespinosa	AMNH 163858	Vietnam, Ha Giang, Yen Minh	KY022210	/
116	B. palpebralespinosa	KIZ 011650	Vietnam, Thanh Hoa, Pu Hu	KX811889	KX812138
117	B. puningensis	SYS a005770	China, Guangdong, Puning	OL635585	OL634853
118	B. puningensis	SYS a007649	China, Guangdong, Puning	OL635589	OL634857
119	B. qianbeiensis	KIZ YPX28756	China, Guizhou, Kuankuoshui	KX811855	KX812120
120	B. qianbeiensis	CIB TZ20190608017	China, Guizhou, Tongzi	MT651554	MT654521
121	B. rubrimera	KIZ 020425	China, Yunnan, Jinping	KX811871	KX812123
122	B. rubrimera	AMS R177676	Vietnam, Lao Cai, Sapa	MF536419	MW086542
123	B. sangzhiensis	SYS a004313	China, Hunan, Sangzhi	MH406802	MH406264
124	B. sangzhiensis	SYS a008386	China, Hunan, Shimen	OQ180982	OQ180870
125	B. sanmingensis	SYS a002499	China, Fujian, Jiangle	MH406699	MH406147
126	B. sanmingensis	SYS a004109	China, Fujian, Shanghang	MW367040	MW365490
127	B. sanmingensis	SYS a002468	China, Guangdong, Mt Fenghuang	MH406693	MH406141
128	B. sanmingensis	SYS a007057	China, Jiangxi, Mt Magu	MW367051	MW365501
129	B. shimentaina	SYS a002822	China, Guangdong, Ruyuan	MK029766	MK029776
130	B. shimentaina	SYS a008309	China, Guangdong, Qujiang	OQ180980	OQ180868
131	B. shimentaina	SYS a002077	China, Guangdong, Yingde	MH406655	MH406092
132	B. shunhuangensis	SYS a002198	China, Guangxi, Lingui	MH406672	MH406112
133	B. shunhuangensis	SYS a002295	China, Guangxi, Mt Mao’er	MH406684	MH406130
134	B. shunhuangensis	HNNU 16SH04	China, Hunan, Mt Shunhuang	MK836027	/
135	B. spinata	KIZ 016100	China, Guizhou, Mt Leigong	KX811864	KX812119
136	B. spinata	SYS a002226	China, Guizhou, Mt Leigong	MH406675	MH406115
137	B. tongboensis	SYS a002484	China, Jiangxi, Mt Tongbo	MH406696	MH406144
138	B. tongboensis	SYS a003227	China, Jiangxi, Mt Tongbo	MH406744	MH406201
139	B. tuberogranulata	KIZ YPX10987	China, Hunan, Sangzhi	KX811823	KX812095
140	B. tuberogranulata	SYS a004310	China, Hunan, Sangzhi	MH406801	MH406263
141	B. wugongensis	SYS a005525	China, Hunan, Mt Dawei	OQ180962	OQ180850
142	B. wugongensis	SYS a005510	China, Hunan, Mt Mufu	OQ180960	OQ180848
143	B. wugongensis	SYS a004801	China, Jiangxi, Anfu	MH406854	MH406316
144	B. wuliangshanensis	KIZ YN0705150	China, Yunnan, Mengla	KX811879	KX812128
145	B. wuliangshanensis	SYS a002983	China, Yunnan, Mt Ailao	MH406730	MH406182
146	B. wuliangshanensis	SYS a003924	China, Yunnan, Mt Wuliang	MH406771	MH406230
147	B. wushanensis	KIZ YPX47799	China, Chongqing, Wushan	KX811835	/
148	B. wushanensis	SYS a003009	China, Hubei, Shennongjia	MH406733	MH406185
149	B. wushanensis	KIZ 045469	China, Sichuan, Mt Guangwu	KX811838	KX812094
150	B. xiangnanensis	SYS a005078	China, Guangxi, Guanyang	MH406874	MH406336
151	B. xiangnanensis	SYS a005062	China, Guangxi, Hezhou	MH406872	MH406334
152	B. xiangnanensis	SYS a002874	China, Hunan, Shuangpai	MH406713	MH406165
153	B. xiangnanensis	SYS a002870	China, Hunan, Lanshan	OQ180957	OQ180845
154	B. xianjuensis	SYS a002668	China, Zhejiang, Mt Dapan	MH406704	MH406154
155	B. xianjuensis	SYS a006399	China, Zhejiang, Mt Longmen	OQ180967	OQ180855
156	B. xianjuensis	CIB XJ190503	China, Zhejiang, Xianju	MN563758	MN563774
157	B. xuefengmontis	SYS a007227	China, Hunan, Mt Xuefeng	OQ180973	OQ180861
158	B. xuefengmontis	SYS a004364	China, Hunan, Wugang	MH406813	MH406275
159	B. xuefengmontis	SYS a004365	China, Hunan, Wugang	MH406814	MH406276
160	B. yangmingensis	SYS a002888	China, Hunan, Shuangpai	MH406719	MH406171
161	B. yangmingensis	SYS a002890	China, Hunan, Shuangpai	MH406721	MH406173
162	B. yaoshanensis	SYS a008800	China, Guangxi, Guiping	OQ180986	OQ180874
163	B. yaoshanensis	KIZ 05198	China, Guangxi, Jinxiu	KX811833	KX812097
164	B. yaoshanensis	SYS a004878	China, Guangxi, Jinxiu	MH406863	MH406325
165	B. yingdeensis	SYS a004721	China, Guangdong, Yingde	MH406846	MH406308
166	B. yingdeensis	SYS a005447	China, Guangdong, Yingde	MH406898	MH406360
167	B. yunkaiensis	SYS a004638	China, Guangdong, Xinyi	MH406844	MH406306
168	B. yunkaiensis	SYS a004694	China, Guangdong, Xinyi	MH406845	MH406307
169	X. mangshanensis	SYS a002177	China, Guangdong, Huaiji	MH406666	MH406106
170	X. glandulosa	SYS a003757	China, Yunnan, Mt Gaoligong	MH406754	MH406213