Environmental variables Mean Minimum Maximum Coefficient of variation (%) Width (m) 3.60 1.15 5.9 139.28 Length (m) 6.05 3.00 11.5 299.75 Area (m2) 21.08 4.83 53.34 57.43 Depth (cm) 12.11 5.50 16.03 26.63 Flow (m/sec) 1.47 0.97 2.19 391.07 Shading (%) 62.65 17.22 100.00 152.82 Sand (%) 0.15 0.00 2.50 287.68 Rock (%) 12.35 0.00 60.00 94.04 Gravel (%) 0.01 0.00 0.14 87.04 Pebble (%) 27.76 0.01 72.50 282.29 Boulder (%) 26.16 0.23 78.33 20.23 Root (%) 0.01 0.00 0.08 39.88