Marine shelled Heterobranchia in part (Gastropoda) of the Miocene Central Paratethys Sea Author Harzhauser, Mathias 0000-0002-4471-6655 Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 mathias.harzhauser@nhm.at Author Landau, Bernard M. 0000-0002-7768-8494 Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 bernardmlandau@gmail.com Author Malaquias, Manuel António E. 0000-0002-4471-6655 Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 mathias.harzhauser@nhm.at text Zootaxa 2026 2026-01-08 5742 1 1 150 https://doi.org/10.11646/zootaxa.5742.1.1 journal article 315392 10.11646/zootaxa.5742.1.1 481d3f36-40cd-41b2-8cad-2972369862ae 1175-5334 19166173 150D7DC5-AE6B-4E55-8223-E05630663C71

TABLE 1. List of species described with stratigraphic distribution in the Central Paratethys Sea.

		Eggenburgian	Ottnangian	Karpatian	early Badenian	middle Badenian	late Badenian	Sarmatian
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1	Acteon antipinguis sp. nov.	0	0	0	1	1	0	0
2	Acteon gutta sp. nov.	0	0	0	0	1	0	0
3	Acteon romanicus sp. nov.	0	0	0	0	1	0	0
4	Acteon semistriatus (Férussac, 1822)	1	1	1	1	1	0	0
5	Acteon laevigatus (Grateloup, 1828)	0	0	0	0	1	0	0
6	Acteon trifasciatus sp. nov.	0	0	0	1	1	0	0
7	Acteon viciani sp. nov.	0	1	0	0	0	0	0
8	Acteon vindobonensis sp. nov.	0	0	0	0	1	0	0
9	Acteon sp.	0	0	0	1	0	0	0
10	Crenilabium reussi (Boettger, 1904)	0	0	0	0	1	0	0
11	Rictaxis austriacus sp. nov.	0	0	0	1	0	0	0
12	Rictaxis siedli sp. nov.	0	0	0	1	0	0	0
13	Rictaxis ursulae sp. nov.	0	0	0	1	0	0	0
14	Ringiculocosta cancellarioides (Seguenza, 1881)	0	0	0	1	1	0	0
15	Ringiculocosta costata (Eichwald, 1830)	0	0	0	0	1	1	0
16	Ringicula daisyae sp. nov.	0	0	0	0	1	0	0
17	Ringicula dentata Berger, 1954	0	0	0	0	1	0	0
18	Ringicula guzhovi Landau et al., 2023	0	0	0	0	1	1	0
19	Ringicula hoernesii Seguenza, 1881	0	0	0	0	1	0	0
20	Ringicula laevigata (Eichwald, 1830)	0	0	1	1	1	1	0
21	Ringicula minor (Grateloup, 1838)	1	1	1	1	1	1	0
22	Candinia krachi (Bałuk & Jakubowski, 1968)	0	0	0	0	0	0	0
23	Bulla? elalbanii sp. nov.	0	0	0	0	1	0	0
24	Bulla hungarica Boettger, 1907	0	0	0	0	1	0	0
25	Pyrunculus elongatus (Eichwald, 1830)	0	0	0	0	1	1	0
26	Pyrunculus frequens (Boettger, 1902)	0	0	0	0	1	0	0
27	Pyrunculus latesulcatus (Boettger, 1902)	0	0	0	0	1	0	0
28	Pyrunculus leuchsi (Berger, 1949)	0	0	0	0	1	1	0
29	Pyrunculus microromanica sp. nov.	0	0	0	0	1	0	0
30	Pyrunculus	0	0	0	0	1	0	0
	parangistoma (Boettger, 1902)
31	Pyrunculus szobiensis sp. nov.	0	0	0	0	1	0	0
32	Pyrunculus zuschini sp. nov.	0	0	0	0	1	0	0
33	Retusa mystica sp. nov.	0	0	0	0	1	1	0
34	Retusa promamillata Boettger, 1907	0	0	0	1	1	1	0
35	Retusa sarmatica Berger, 1953	0	0	0	0	1	1	1
36	Retusa sinzovi (Kolesnikov, 1935)	0	0	0	0	0	0	1
37	Retusa truncatula (Bruguière, 1792)	0	0	1	1	1	1	1
38	Retusa vindobonensis (Berger, 1949)	0	0	0	0	1	0	0
39	Retusa voeslauensis (Berger, 1953)	0	0	0	0	1	1	0
40	Volvulella acuminata (Bruguière, 1792)	0	0	0	0	1	0	0
41	Acteocina fusiformis (Boettger, 1907)	0	0	1	1	1	1	0
42	Acteocina lichtensteinii (Eichwald, 1830)	0	0	0	0	0	1	0
43	Acteocina melinda (De Gregorio, 1885)	0	0	0	0	0	0	1
44	Acteocina volhynica (Eichwald, 1830)	0	0	0	0	0	0	1
45	Acteocina sp.	0	0	0	0	0	0	1
46	Acrostemma edlaueri (Berger, 1953)	0	0	0	0	1	0	0
47	Acrostemma kuehnelti (Berger, 1953)	0	0	0	0	1	0	0
48	Cylichna caudex sp. nov.	0	0	1	1	1	0	0
49	Cylichna cylindracea	0	0	0	0	1	0	0
	(Pennant, 1777)
50	Cylichna dolium sp. nov.	0	0	0	0	1	0	0
51	Cylichna eschnerae sp. nov.	0	0	0	1	0	0	0
52	Cylichna intermedia (Sacco, 1897)	0	0	1	1	1	1	0
53	Cylichna miosimplex sp. nov.	0	0	0	0	1	0	0
54	Cylichna steiningeri sp. nov.	1	0	0	0	0	0	0
55	Cylichna straussi sp. nov.	1	0	0	0	0	0	0
56	Cylichna subbacillus Berger. 1953	0	0	0	0	1	0	0
57	Cylichna sp. 1	0	0	0	0	1	0	0
58	Cylichnella bitaeniata (Boettger, 1902)	0	0	0	0	1	1	0
59	Neubaueriella lapugyensis (Berger, 1949)	0	0	0	0	1	0	0
60	Neubaueriella sp.	0	0	1	0	0	0	0
61	Haminoea miocaenica	0	0	0	0	1	0	0
	(Berger, 1949)
62	Vellicolla pannonica (Csepreghy-Meznerics, 1952)	0	0	0	0	1	0	0
63	Vellicolla postneglecta (Berger, 1953)	0	0	0	0	1	0	0
64	Melanochlamys radwanskii (Bałuk, 2018)	0	0	0	0	1	0	0
65	Chelidonura piseraii sp. nov.	0	0	0	0	1	0	0
66	Roxania bomba nov. nom.	0	0	0	1	0	0	0
67	Roxania callifera (Boettger, 1907)	0	0	0	0	1	0	0
68	Roxania oviformis sp. nov.	0	0	0	0	1	0	0
69	Roxania pila sp. nov.	0	0	0	0	1	0	0
70	Roxania utriculus (Brocchi, 1814)	0	0	1	1	1	0	0
71	Laona quadrata (Wood, 1839)	0	0	0	0	1	0	0
72	Cosmophiline marinellii Berger, 1949	0	0	0	0	1	0	0
73	Hermania scabra (Müller, 1784)	0	0	0	0	1	0	0
74	Philine catena (Montagu, 1803)	0	0	0	0	1	0	0
75	Sabatia quasimodoi sp. nov.	0	0	0	0	1	0	0
76	Scaphander alatissimus sp. nov.	0	0	0	1	0	0	0
77	Scaphander grundincola (De Gregorio, 1885)	0	0	0	1	1	1	0
78	Scaphander helveticus Peyrot, 1932	0	0	1	0	0	0	0
79	Scaphander catena sp. nov.	0	0	0	0	1	0	0
80	Scaphander perforatus sp. nov.	0	0	0	0	1	0	0
81	Scaphander quadratus	0	0	0	0	1	0	0
	sp. nov.
82	Scaphander schollnbergeri sp. nov.	0	0	0	0	1	0	0
83	Pseudavena bella sp. nov.	0	0	0	0	1	0	0
	total	4	3	10	20	65	16	6