Amphibian and reptilian fauna from the early Miocene of Echzell, Germany Author Vasilyan, Davit https://orcid.org/0000-0001-8712-0678 JURASSICA Museum, Route de Fontenais 21. 2900 Porrentruy, Switzerland & Department of Geosciences, University of Fribourg, Chemin du musee 6, 1700, Fribourg, Switzerland davit.vasilyan@jurassica.ch Author Cernansky, Andrej https://orcid.org/0000-0001-8920-2503 Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynska dolina, Ilkovicova 6, 842 15 Bratislava, Slovakia Author Szyndlar, Zbigniew Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Slawkowska 17, 31 - 016 Krakow, Poland Author Moers, Thomas https://orcid.org/0000-0003-2268-5824 Department of Palaeobiology, Swedish Museum of Natural History, P. O. Box 50007, 104 05 Stockholm, Sweden & Bolin Centre for Climate Research, Stockholm University, Stockholm, Sweden text Fossil Record 2022 2022-05-10 25 1 99 145 http://dx.doi.org/10.3897/fr.25.83781 journal article http://dx.doi.org/10.3897/fr.25.83781 2193-0074-1-99 7A16698D4F1848D29D9651A6E0CC15AC 2F5D6AE2EEB55A17ACF1623B06B4EA8D

Table 2. Overview of amphibian and reptilian faunas from MN3-MN4 sites. Each large taxonomic group is highlighted in different color. The herpetofaunistic record follows for: Wintershof West - Boehme and Ilg (2003); Ahnikov 1 (= Merkur North) - Cernansky (2010a), Cernansky and Bauer (2010), Cernansky et al. (2015), Klembara et al. (2010), Villa and Delfino (2019a, b); Oberdorf - Cernansky (2016), Cernansky et al. (2020b), Sanchiz (1998), Szyndlar (1998), Villa and Delfino (2019a); Petersbuch 2 - Cernansky (2011), Ivanov (2001), Klembara et al. (2010), Szyndlar and Schleich (1993), Szyndlar and Rage (2002), Szyndlar and Rage (2003), Villa and Delfino (2019a), Villa et al. (2018); Echzell - present study; Mokra-W Q 1/2001 and Mokra-W Q 2/ 2001 - Ivanov (2008), Ivanov et al. (2020); Dolnice - Boehme and Ilg (2003), Cernansky (2010b), Fejfar and Rocek (1986), Klembara (2015), Szyndlar (1987), Villa and Delfino (2019a); Erkertshofen 1 - Klembara et al. (2010); Beon 1 - Rage and Bailon (2005); Rembach - Klembara et al. (2010); Forsthart - Klembara et al. (2010). The ecophysiological indices for Echzell assemblage are given for each taxon as well as the calculated precipitation values and errors are calculated according to Boehme et al. (2006).

		taxon	Wintershof West	Ahnikov 1 (=Merkur North)	Oberdorf	Petersbuch 2	Echzell	Mokra-W Q 1/2001	Mokra-W Q 2/2001	Dolnice	Erkertshofen 1	Beon 1	Rembach	Forsthart
			MN3a	MN 3a	MN4	MN4a	MN4	MN4	MN4	MN 4b	MN 4b	MN 4b	MN 4b	MN 4b
Allocaudata	Albanerpetonidae	Albanerpeton inexpectatum	+	cf.	+	+	0.0917				+
Urodela	Cryptobranchidae	Andrias scheuchzeri		+
	Proteidae	Mioproteus sp.							+
	Salamandridae	Salamandrina sp.				+
		Chioglossa meini				+			+		+
		Chioglossa sp.			+		0.9768
		Mertensiella mera							+
		Mertensiella sp.	+	+		+	0.9768		+
		Salamandra sansaniensis	+		+	+					+
		Salamandra sp.					0.3918
		Chelotriton paradoxus	+			+
		Chelotriton aff. pliocaenicus				+
		Chelotriton sp.	+	+	+		0.3918				+	cf.
		Chelotriton sp. 1						+	+
		Chelotriton sp. 2						+	+
		Triturus cf. marmoratus						+	+			+
		Triturus ex. gr. cristatus							+
		Triturus (Triturus marmoratus group) sp.				+
		Lissotriton cf. helveticus				+						+
		Lissotriton roehsi			+			+	+
		Lissotriton aff. roehsi						+
		Lissotriton sp.					0.3918
		Triturus / Lissotriton sp.		+
		Triturus aff. vittatus				+
		Ichthyosaura alpestris
		Ichthyosaura wintershofi	+
Anura	Alytidae	Latonia gigantea								+
		Latonia ragei	+	+	+	+
		Latonia cf. ragei	+									+
		Latonia sp.					0.3918
	Palaeobatrachidae	Palaeobatrachus robustus					1
		Palaeobatrachus sp.		+	+
		Palaeobatrachidae sp. 1				+
		Palaeobatrachidae sp. 2				+
	Bufonidae	Bufo sp.							+
		Bufotes (cf.) Bufotes viridis				+					+
	Ranidae	Pelophylax sync esculenta						+	+
		Pelophylax sp.		+	+	+	0.513			+	+	+
		Rana sp.	+				0.3918							+
	Pelodytidae	Pelodytes sp.	+	+	+						+
	Pelobatidae	Eopelobates sp.	+	+						+
		Pelobates sanchizi			+		0.0917	+	+
		Pelobates sp.			+	+					+			+
		Pelobates sp. nov.				+
	Hylidae	Hyla sp.			+					+
Crocodylia		Diplocynodon sp.		+						+		+
Testudines		Ptychogaster sp.	+			+				+	+	+
		Ptychogaster buechelbergense		+						cf.
		Testudo rectangularis												+
		Testudo sp.				+				+	+	+		+
		Geochelone / Ergilemys sp.				+						+
		aff. Clemmydopsis sp.											+
Serpentes	Colubridae	Natrix longivertebrata					+					cf.
		Natrix merkurensis		+								aff.
Serpentes	Colubridae	Natrix sansaniensis		+		+						+
		Natrix sp.						+	+
		cf. Neonatrix sp.			+	+
		Palaeonatrix lehmani				+				+		aff.
		Neonatrix nova								+
		Neonatrix europaea										+
		Neonatrix natricoides										+
		Palaeonatrix sp.			+
		Natricinae indet.		+		+		+	+
		Coluber caspioides		+	cf.	+
		Coluber dolnicensis		+						+
		Coluber pouchetii										+
		Coluber suevicus		+
		Coluber sp.						+	+
		Texasophis meini										+
		Texasophis bohemicus								+
		Elaphe sp.		+
		Colubrinae indet.		+			+
		Colubrinae indet. 1						+	+
		Colubrinae indet.	+						+
		Colubrinae indet. A			+
		Colubrinae indet. B			+
		Colubridae indet.		+				+	+
		Colubroidea indet.					+
	Viperidae	Vipera antiqua				+				+
		Vipera platyspondyla				+				+
		Vipera aspis complex										+
		Daboia /oriental Vipera complex										+
		Viper sp.		+	+				+
		Viperidae indet.						+	+
		Viperinae indet.						+	+
	Elapidae	Naja romani				+
		Naja cf. romani					+
		Micrurus gallicus				+						+
		Elapidae indet.		+				+				+
		Elapidae / Natricinae indet.			+
	Boidae	Bavarioboa hermi				+		+		+
		Bavarioboa cf. hermi					0.0917	+
		Bavarioboa sp.		+
		cf. Bavarioboa sp.			+
		Pyton europaeus										+
		Pyton sp.						+
		Falseryx petersbuchi				+
		Bransateryx septentrionalis				+				+
		Bransateryx sp.
		Scolecophidia indet.										+
	Aniliidae	Eoanilus sp.				+
		Serpentes indet.	+	+
Iguania	Chamaeleonidae	Chamaeleo sp.				+							+
		Chamaeleo andrusovi					0.0917			+
		Chamaeleo caroliquarti	+							+	+
		Chamaeleonidae indet.		+	?	+	+
Gekkota	Gekkonidae	Euleptes gallica		+
		Gekkota indet.	+	+	+		0.0917					+	+
Amphisbaenia	Blanidae	Blanus sp.								+			+
		Blanus thomaskelleri		+
	Amphisbaenidae	Amphisbaenidae indet.									+	+
		Amphisbaenia indet.							+
Lacertiformata	Lacertidae	Lacerta cf. L. poncenatensis			+
		Lacerta ex. gr. L. viridis								+
		Lacerta sp.		+	+	+				+	+		+
		Lacerta sp. 1	+											+
		Lacerta sp. 2	+											+
		Miolacerta tenuis								+
		Miolacerta sp.				+							+
		Janoskia sp.				+
		Ambyolacerta dolnicensis								+
Lacertiformata	Lacertidae	Ambyolacerta sp.									+
		Lacertidae indet.		+	+		0	+	+	+		+		+
		Lacertidae indet. 1										+
		Lacertidae indet. 2										+
		Lacertoidea indet.							+
Scinciformata	Scincidae	Eumeces sp.									+
		Bavariascincus mabuyaformis				+							+	+
		Chalcides augei			+
		cf. Chalcides sp.					0
		Scincidae indet.				+	+			+	+
	Cordylidae	Palaeocodylus bohemicus								+
		aff. Palaeocodylus bohemicus		+
		Cordylidae indet.		?	+	+
		Scincoidea indet.			+				+
Anguimorpha	Varanidae	Varanus mokrensis						+	+
		Varanus hofmanni				+
		Varanus sp.									+	+
		cf. Varanus Varanus sp.				+
	Shinisauridae	Merkurosaurus ornatus		+
		Shinisauridae indet.	+
	Anguidae	Pseudopus laurillardi				+		+	+		+	+
		Pseudopus confertus		+
		Pseudopus ahnikoviensis		+						+
		Pseudopus sp.	+					+	+	+			+	+
		Smithosaurus echzellensis					+
		Anguis sp.								+
		Ophisaurus fejfari				+				+
		Ophisaurus cf. O. fejfari	+
		Ophisaurus spinari		+						+
		Ophisaurus cf. O. spinari			+
		Ophisaurus robustus								+
		Ophisaurus aff. O. robustus									+
		Ophisaurus holeci		+						+
		Ophisaurus sp.		+	+	+		+		+		+		+
		Ophisaurus sp. 1		+						+			+
		Ophisaurus sp. 2		+						+			+
		Anguinae indet.					+	+	+
		Anguidae indet.			+	+	+						+
		Squamata indet.					+
		Squamata indet. 1			+
		Squamata indet. 2			+
		Squamata indet. 3			+
		average ecophysiological index					0.344
		MAP					791
		recent MAP (in mm)					630
		95 % prediction interval - error (in mm)					254
		(near) city/town of the value or recent MAP			Frankfurt a. M.
		MAP/MAPrecent (in %)			125.5036157