Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands Author Archibald, S. Bruce 0000-0002-4397-2497 Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada bruce.archibald@ubc.ca Author Rasnitsyn, Alexandr P. A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca text Zootaxa 2023 2023-05-04 5278 1 1 38 http://dx.doi.org/10.11646/zootaxa.5278.1.1 journal article 54646 10.11646/zootaxa.5278.1.1 1528757a-5499-40b9-a728-c5bf0c770b27 1175-5326 7894826 BE0A07DB-170A-4B72-8178-9B8144C1FBF6

TABLE 1. Fossil record of Cimbicidae with selected references.

Paleogene
Paleocene
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Selandian
Menat Formation, France
Cenocimbicinae
Cenocimbex menatensis Nel, 2004	Nel (2004)
Eocene
Ypresian
Okanagan Highlands, Canada and USA
Cenocimbicinae
Allenbycimbex morrisae n. gen. and sp.	herein
Leptostigma alaemacula n. gen. and sp.	herein
Leptostigma brevilatum n. gen. and sp.	herein
Leptostigma fasciatum n. gen. and sp.	herein
Leptostigma longiclava n. gen. and sp.	herein
Leptostigma longipallidum n. gen. and sp.	herein
Leptostigma longitenebricum n. gen. and sp.	herein
Leptostigma proxivena n. gen. and sp.	herein
Subfamily incertae sedis
Genus and species incertae sedis	herein
Green River Formation, Colorado, USA
Pachylostictinae
Eopachylosticta byrami (Cockerell, 1925)	Cockerell (1925), Malaise (1945)
Priabonian
Florissant Formation, United States of America
Abiinae
Trichiosomites obliviosus Brues, 1908	Brues (1908)
Cimbicinae
Floricimbex vetusculus (Cockerell, 1922)	Cockerell (1922)
Phenacoperga coloradensis (Cockerell, 1907)	Cockerell (1907, 1908), Rohwer (1908)
Baltic amber, Russia
Subfamily incertae sedis
Cimbex sp. (larva)	Menge (1856), Handlirsch (1907), APR pers. obs.
Neogene
Miocene
Aquitanian/Burdigalian
Masaragawa Formation, Japan
Cimbicinae
? Cimbex sp.	Fujiyama (1985)
Burdigalian/Langhian
Shanwang Formation, China
Cimbicinae
Cimbex chromoptera * Zhang, 1989	Zhang (1989)
Pseudoclavellaria autochthona (Zhang)	Zhang (1989), Taeger et al. (2010)
Pseudoclavellaria bicolor Zhang, 1989	Zhang (1989)