Eastern Colombian Paramo Liodessus Guignot, 1939 diving beetles are genetically structured, but show signs of hybridization, with description of new species and subspecies (Coleoptera, Dytiscidae) Author Balke, Michael https://orcid.org/0000-0002-3773-6586 SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany & GeoBioCenter, Ludwig Maximilians University, Munich, Germany balke.m@snsb.de Author Neven, Katja SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany Author Villastrigo, Adrian https://orcid.org/0000-0002-3531-0821 SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany Author Ospina-Torres, Rodulfo Departamento de Biologia, Universidad Nacional de Colombia-Sede Bogota, Bogota, Colombia Author Prieto, Carlos Departamento de Biologia, Universidad del Atlantico, Barranquilla, Colombia & Corporacion Universitaria Autonoma del Cauca, Popayan, Colombia Author Gutierrez Rubiano, Nicolas Departamento de Biologia, Universidad Nacional de Colombia-Sede Bogota, Bogota, Colombia Author Lotta, Ingrid Departamento de Biologia, Universidad Nacional de Colombia-Sede Bogota, Bogota, Colombia Author Duenas, Luisa F. https://orcid.org/0000-0002-0756-826X Departamento de Biologia, Universidad Nacional de Colombia-Sede Bogota, Bogota, Colombia Author Hendrich, Lars https://orcid.org/0000-0001-8366-0749 SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany text ZooKeys 2023 2023-01-31 1143 165 187 http://dx.doi.org/10.3897/zookeys.1143.97461 journal article http://dx.doi.org/10.3897/zookeys.1143.97461 1313-2970-1143-165 53A716C7B6C24DB6A1CFEE5D98FE5193 494000F0E813503D8B8C25F5A4C91BC5

Table 1. Diagnostic characters in the Cox1 alignment of the COLLI project on BOLD webportal. Above two taxa relative to all other entries in COLLI, and below, within the L. bogotensis complex. Numbers above indicate nucleotide position in the 658 bp alignment.

Taxon, Number of sequences	19	22	25	49	58	59	82	88	91	106	127	206	217	223	232	256	286	290	304	316	325
Liodessus santarosita (6)	C		T	C	T				A			C		T	A	C	T	A
Liodessus bogotensis (55)
L. bogotensis almorzadero (4)			G					A
L. bogotensis bogotensis (21)						T
L. bogotensis chingaza (16)		A								T	C
L. bogotensis lacunaviridis (4)							C												A	A	G
L. bogotensis matarredonda (5)													G
L. bogotensis sumapaz (5)
	331	334	337	352	355	358	412	421	445	472	473	490	511	542	547	548	553	586	616	643	655
Liodessus santarosita		G	T	A		C			C								G	A		T
Liodessus bogotensis					T										C
L. bogotensis almorzadero														T
L. bogotensis bogotensis
L. bogotensis chingaza										G		C									G
L. bogotensis lacunaviridis	G						G	C								T			T
L. bogotensis matarredonda
L. bogotensis sumapaz											A		G