Euroscaptor darwini sp. nov., a new species of mole (Mammalia, Eulipotyphla, Talpidae) from the north-central mountains in Vietnam Author Nguyen, Son Truong 0000-0003-3214-4407 Institute of Biology, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet Road, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet Road, Hanoi, Vietnam Author Bui, Hai Tuan 0000-0003-4065-7229 Faculty of Biology, Hanoi University of Science, Vietnam National University (VNU), 334 Nguyen Trai Road, Hanoi, Vietnam Author Dau, Vinh Quang 0000-0002-8976-1644 Hong Duc University, 565 Quang Trung Road, Thanh Hoa, Vietnam Author Le, Phuong Dinh Pu Luong Nature Reserve, Thanh Hoa, Vietnam Author Vu, Yen Huong https://orcid.org/0009-0001-4643-167X Institute of Biology, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet Road, Hanoi, Vietnam & Faculty of Biology, Hanoi University of Science, Vietnam National University (VNU), 334 Nguyen Trai Road, Hanoi, Vietnam text ZooKeys 2025 2025-10-10 1255 239 274 journal article 312575 10.3897/zookeys.1255.161942 1abe6cfb-401a-4d86-94e7-6b3023e65ed2 3C3E0CF2-85BF-4F5C-B4D3-9140F0BE2BC9

Table 5. Craniodental measurements (in mm) of type specimens of Euroscaptor darwini sp. nov. (Voucher: 1 ♂ NTS.2024.PL.01; 4 ♀♀ NTS.2025.PL.02 – NTS.2025.PL.05) and E. subanura (Voucher: 4 ♂♂ SIK 0882, SIK 0884, SIK 0875, SIK 0876; 1 ♀ SIK 0883), as well as additional specimens representing three other Vietnamese populations of E. subanura from Pu Huong NR (Nghe An Province), Na Hang (Tuyen Quang Province), and Xuan Son NP (Phu Tho Province).

Character	E. darwini sp. nov.		E. subanura		E. subanura	E. subanura	E. subanura
	Pu Luong NR (Thanh Hoa) pop.		Tam Dao (Tuyen Quang) pop.		Pu Huong NR (Nghe An) pop.	Na Hang (Tuyen Quang) pop.	Xuan Son NP (Phu Tho) pop.
	Male (1)	Female (4)	Male (4)	Female (1)	Female (1)	Male (2)	Male (3)	Female (6)
GLS	30.51	30.94–31.01 30.97 ± 0.03	30.44–30.96 30.66 ± 0.22	31.41	31.33	30.75–31.69 31.22 ± 0.66	30.49–30.98 30.78 ± 0.26	29.96–31.02 30.48 ± 0.57
BB	13.76	13.84–14.26 14.08 ± 0.21	13.44–14.23 13.75 ± 0.34	14.44	14.33	13.93–14.67 14.30 ± 0.52	13.91–14.47 14.15 ± 0.29	13.58–14.19 13.92 ± 0.23
IOB	6.72	6.83–6.87 6.85 ± 0.02	6.57–6.86 6.77 ± 0.14	6.97	6.74	7.13–7.14 7.14 ± 0.01	6.59–7.05 6.79 ± 0.24	6.53–7.13 6.82 ± 0.21
BIOF	5.41	5.59–5.77 5.68 ± 0.09	6.13–6.21 6.17 ± 0.04	6.21	6.27	6.27–6.35 6.31 ± 0.06	6.12–6.22 6.16 ± 0.05	6.16–6.38 6.29 ± 0.08
RB	4.12	4.17–4.27 4.21 ± 0.04	3.84–4.29 4.10 ± 0.19	4.21	4.26	3.99–4.41 4.20 ± 0.3	3.96–4.16 4.03 ± 0.11	4.04–4.28 4.13 ± 0.09
LZA	8.22	7.88–8.02 7.96 ± 0.06	7.24–7.61 7.43 ± 0.16	7.68	7.63	7.01–7.54 7.28 ± 0.37	7.15–7.51 7.28 ± 0.2	7.11–7.51 7.28 ± 0.17
PZRW	10.23	10.09–10.37 10.21 ± 0.13	10.01–10.17 10.09 ± 0.08	10.13	9.85	9.96–10.18 10.07 ± 0.16	9.19–9.64 9.44 ± 0.23	9.34–10.16 9.78 ± 0.36
ACRB	6.92	6.98–7.19 7.09 ± 0.11	7.12–7.26 7.19 ± 0.07	7.29	7.18	7.27–7.35 7.31 ± 0.06	7.01–7.16 7.11 ± 0.08	7.11–7.28 7.21 ± 0.07
FMW	3.35	3.38–3.54 3.45 ± 0.07	3.27–3.71 3.46 ± 0.19	3.27	3.67	3.35–3.58 3.47 ± 0.16	3.51–3.67 3.57 ± 0.09	3.45–3.75 3.63 ± 0.13
CCoL	21.41	21.54–21.88 21.64 ± 0.16	21.42–21.83 21.68 ± 0.2	22.31	22.26	21.49–22.13 21.81 ± 0.45	20.92–21.48 21.05 ± 0.45	21.15–21.91 21.52 ± 0.42
PL	11.61	11.81–12.16 12 ± 0.17	11.55–12.53 12.03 ± 0.4	12.6	12.32	11.72–12.51 12.12 ± 0.56	11.98–12.38 12.12 ± 0.22	12.11–12.69 12.38 ± 0.31
I 1 – M 3	11.51	11.59–12.02 11.81 ± 0.22	11.86–12.14 11.98 ± 0.12	12.72	12.16	11.89–12.33 12.11 ± 0.31	11.43–12.02 11.67 ± 0.31	11.58–12.21 11.84 ± 0.35
I 1 – P 4	7.15	7.23–7.53 7.38 ± 0.17	7.17–7.36 7.25 ± 0.08	7.79	7.27	7.53–7.58 7.56 ± 0.04	6.73–7.68 7.21 ± 0.39	7.23–7.44 7.33 ± 0.11
I 1 – P 3	5.92	5.72–6.07 5.92 ± 0.16	5.56–5.88 5.69 ± 0.16	6.03	5.78	5.95–6.06 6.01 ± 0.08	5.16–5.97 5.61 ± 0.35	5.68–5.95 5.85 ± 0.15
I 1 – C	3.12	2.94–3.17 3.01 ± 0.11	2.62–3.04 2.84 ± 0.17	2.92	2.99	2.97–3.09 3.03 ± 0.08	2.76–3.09 2.96 ± 0.17	2.68–3.09 2.88 ± 0.17
C – M 3	10.13	10.14–10.44 10.28 ± 0.15	10.47–10.76 10.58 ± 0.13	11.21	10.55	10.24–10.69 10.47 ± 0.32	10.03–10.55 10.24 ± 0.28	9.76–10.76 10.17 ± 0.44
C – P 4	5.51	5.56–5.93 5.72 ± 0.21	5.56–5.71 5.65 ± 0.07	6.27	5.49	5.63–5.79 5.71 ± 0.11	5.81–6.08 5.95 ± 0.14	5.31–6.05 5.70 ± 0.37
P 4 – M 3	5.91	6.12–6.18 6.15 ± 0.03	6.27–6.41 6.32 ± 0.06	6.58	6.39	6.28–6.45 6.37 ± 0.12	5.81–6.11 5.92 ± 0.17	5.71–6.26 5.99 ± 0.2
P 1 – P 4	3.99	4.02–4.11 4.07 ± 0.05	4.19–4.23 4.21 ± 0.02	4.27	4.25	4.16–4.25 4.21 ± 0.06	4.18–4.29 4.23 ± 0.06	4.13–4.29 4.21 ± 0.07
M 1 – M 3	4.54	4.65–5.03 4.80 ± 0.17	4.81–4.98 4.92 ± 0.08	5.21	4.91	4.75–5.01 4.88 ± 0.18	4.43–4.71 4.54 ± 0.15	4.34–4.87 4.55 ± 0.23
M 2 – M 2 W	6.69	6.72–7.17 7.02 ± 0.2	7.01–7.33 7.12 ± 0.14	7.81	7.14	7.02–7.26 7.14 ± 0.17	6.71–7.06 6.86 ± 0.18	6.65–7.29 6.96 ± 0.22
CCW	3.71	3.64–3.85 3.75 ± 0.1	3.57–3.71 3.64 ± 0.07	3.73	3.74	3.71–3.76 3.74 ± 0.04	3.49–3.61 3.55 ± 0.06	3.41–3.79 3.56 ± 0.13
PWCC	2.41	2.27–2.42 2.35 ± 0.06	2.17–2.37 2.31 ± 0.09	2.41	2.54	2.31–2.36 2.34 ± 0.04	2.29–2.33 2.31 ± 0.02	2.16–2.47 2.29 ± 0.13
PWM 2 M 2	3.91	3.81–3.93 3.90 ± 0.06	3.57–3.96 3.80 ± 0.16	4.15	3.95	3.95–3.99 3.97 ± 0.03	3.69–3.89 3.80 ± 0.1	3.56–3.94 3.83 ± 0.14
PWM 3 M 3	3.91	3.92–4.06 3.99 ± 0.06	3.92–4.09 4.03 ± 0.07	4.35	4.11	4.07–4.21 4.14 ± 0.1	3.67–4.19 3.98 ± 0.27	3.91–4.17 4.07 ± 0.12
ML	18.93	19.21–19.37 19.26 ± 0.07	19.1–19.47 19.32 ± 0.17	20.28	19.77	19.07–20.18 19.63 ± 0.78	18.33–19.86 18.89 ± 0.55	18.91–19.69 19.35 ± 0.4
CAL	8.14	8.09–8.24 8.15 ± 0.07	8.05–8.33 8.15 ± 0.12	8.33	8.29	7.81–8.29 8.05 ± 0.34	7.69–8.36 8.05 ± 0.27	8.16–8.41 8.31 ± 0.13
MH	5.84	5.73–5.83 5.76 ± 0.05	5.72–6.01 5.81 ± 0.13	6.03	6.26	5.64–6.08 5.86 ± 0.31	5.53–6.07 5.76 ± 0.19	5.78–6.31 6.00 ± 0.28
i 1 – m 3	10.88	11.01–11.45 11.23 ± 0.19	10.69–11.57 11.18 ± 0.37	11.87	11.67	11.38–11.59 11.49 ± 0.15	11.05–11.33 11.19 ± 0.14	10.51–11.65 10.95 ± 0.46
p 1 – m 3	8.99	9.16–9.43 9.23 ± 0.21	9.24–9.39 9.32 ± 0.08	9.76	9.56	9.21–9.52 9.37 ± 0.22	8.63–9.54 8.99 ± 0.34	9.03–9.31 9.17 ± 0.14
p 4 – m 3	6.07	6.22–6.47 6.33 ± 0.13	6.39–6.54 6.46 ± 0.07	6.67	6.68	6.26–6.48 6.37 ± 0.16	5.91–6.28 6.14 ± 0.2	5.85–6.49 6.13 ± 0.25
i 1 – p 4	6.11	6.21–6.27 6.24 ± 0.03	6.22–6.29 6.25 ± 0.03	6.24	6.19	6.32–6.37 6.35 ± 0.04	6.16–6.24 6.21 ± 0.04	6.17–6.32 6.21 ± 0.06
m 1 – m 3	4.89	4.97–5.29 5.09 ± 0.15	5.15–5.25 5.20 ± 0.04	5.34	5.49	5.09–5.29 5.19 ± 0.14	4.68–5.06 4.84 ± 0.2	4.62–5.28 4.84 ± 0.25
i 1 – c	1.54	1.56–1.63 1.60 ± 0.03	1.72–1.81 1.74 ± 0.09	1.86	1.88	1.77–1.79 1.78 ± 0.01	1.68–1.86 1.74 ± 0.1	1.69–1.88 1.77 ± 0.07
i 1 – p 1	2.98	2.95–3.19 3.06 ± 0.1	3.05–3.17 3.10 ± 0.05	3.02	3.22	3.34–3.39 3.37 ± 0.04	2.77–3.22 3.05 ± 0.18	3.11–3.17 3.13 ± 0.03
p 1 – p 4	3.97	4.07–4.19 4.10 ± 0.1	3.87–4.12 3.99 ± 0.11	4.15	3.94	4.08–4.12 4.10 ± 0.03	3.79–4.23 3.98 ± 0.16	3.93–4.25 4.1 ± 0.16