Morphological and molecular variation reveal cryptic diversity in the racer Philodryas patagoniensis (Girard, 1858) (Squamata: Colubridae) Author Di Pietro, Diego Omar 0000-0001-7329-6429 Sección Herpetología, División Zoología Vertebrados, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, Avenida 122 y 60 s / No, 1900, La Plata, Buenos Aires, Argentina & Consejo Nacional de Investigaciones Científicas y Técnicas-Centro Científico Tecnológico La Plata, Calle 8 No 1467, 1900, La Plata, Buenos Aires, Argentina Author Sánchez, Julieta 0000-0003-1192-7613 Laboratorio de Ecología Molecular, Centro Austral de Investigaciones Científicas (CADIC-CONICET), Calle Bernardo A. Houssay No 200, 9410, Ushuaia, Tierra del Fuego, Argentina Author Poljak, Sebastián 0000-0001-7972-9352 Laboratorio de Ecología Molecular, Centro Austral de Investigaciones Científicas (CADIC-CONICET), Calle Bernardo A. Houssay No 200, 9410, Ushuaia, Tierra del Fuego, Argentina & Instituto de Ciencias Polares, Ambiente y Recursos Naturales, Universidad Nacional de Tierra del Fuego, Fuegia Basket No 251, 9410, Ushuaia, Tierra del Fuego, Argentina Author Alcalde, Leandro 0000-0002-4365-2434 Consejo Nacional de Investigaciones Científicas y Técnicas-Centro Científico Tecnológico La Plata, Calle 8 No 1467, 1900, La Plata, Buenos Aires, Argentina & Sección Herpetología, Instituto de Limnología Dr. R. A. Ringuelet (CONICET-UNLP), Boulevard 120 y 62 s / No, 1900, La Plata, Buenos Aires, Argentina text Vertebrate Zoology 2026 2026-02-10 76 93 119 journal article 314208 10.3897/vz.76.e169219 b1b36b42-6c45-433c-91b1-354faac1ae1a F5504E2E-AFE9-4EE6-AE74-8AC52746A1E3
Global pholidosis variation of each morphotype of P. patagoniensis. The most common formula for each pholidosis type is highlighted in bold.
Pholidosis typeA – morphotypeB – morphotype
Preoculars1 (n = 41) – 1 / 2 (n = 1)1 (n = 99)
Postoculars2 (n = 42)2 (n = 99)
Temporals1 + 2 (n = 32) – 1 + 2 + 3 / 1 + 2 (n = 2) 1 + 1 + 2 (n = 2) – 1 + 1 + 2 / 1 + 2 (n = 1) 1 + 2 / 1 + 2 + 3 (n = 1) – 1 + 1 / 1 + 2 (n = 1) 2 + 2 / 1 + 2 (n = 1) – 1 + 2 / 1 + 4 (n = 1) 1 + 3 + 1 / 1 + 2 (n = 1)1 + 2 (n = 70) – 1 + 1 + 2 / 1 + 2 (n = 4) 1 + 2 / 1 + 3 (n = 3) – 1 + 2 + 3 / 1 + 2 (n = 2) 2 + 2 (n = 2) – 2 + 3 (n = 2) 1 + 2 + 3 (n = 1) – 1 + 2 / 2 + 2 (n = 1) 1 + 3 / 1 + 2 (n = 1) – 1 + 1 + 3 / 1 + 3 (n = 1) 1 + 2 / 1 + 1 + 2 (n = 1) – 1 + 2 + 2 / 1 + 2 (n = 1) 1 + 2 + 2 / 1 + 2 + 3 (n = 1) – 1 + 3 + 3 / 2 + 2 + 3 (n = 1) 2 + 3 / 1 + 3 (n = 1) – 1 + 1 / 1 + 1 + 2 (n = 1) 2 + 3 / 2 + 1 + 2 (n = 1) – 1 + 1 / 1 + 2 (n = 1) 1 + 1 + 1 / 1 + 2 (n = 1) – 1 + 2 / 1 + 4 (n = 1) 2 + 1 + 2 (n = 1) – 1 + 2 / 2 + 1 + 2 (n = 1)
Supralabials7 [3, 4] (n = 42)7 [3, 4] (n = 85) – 8 [4, 5] (n = 7) 8 [4, 5] / 7 [3, 4] (n = 3) – 7 [3, 4, 5] / 7 [3, 4] (n = 1) 8 [3, 4, 5] / 8 [4, 5] (n = 1) – 8 [3, 4, 5] / 7 [3, 4] (n = 1) 8 [3, 4] (n = 1)
Infralabials9 [5] (n = 40) – 9 [5] / 8 [5] (n = 1) 11 [6] / 10 [6] (n = 1)9 [5] (n = 81) – 10 [5] / 9 [5] (n = 4) 10 [5] (n = 4) – 9 [5] / 8 [5] (n = 1) 11 [5] (n = 1) – 10 [6] / 9 [5] (n = 1) 8 [5] / 9 [5] (n = 1) – 9 [5] / 10 [5] (n = 1) 9 [6] / 8 [5] (n = 1) – 9 [6] / 9 [5] (n = 1) 10 [6] (n = 1) – 9 [5] / 10 [6] (n = 1)
Dorsals19–19 – 15 (n = 31) – 20–19 – 15 (n = 4) 18–19 – 15 (n = 3) – 19–18 – 15 (n = 2) 19–19 – 16 (n = 1) – 19–15 – 15 (n = 1)19–19 – 15 (n = 89) – 19–19 – 16 (n = 5) 19–18 – 15 (n = 3) – 20–19 – 15 (n = 3) 19–19 – 13 (n = 1) – 20–15 – 15 (n = 1) 21–19 – 15 (n = 1) – 18–19 – 15 (n = 1)