TABLE 5

TABLE 5. Morphological comparison of the subspecies of *Eisenia nordenskioldi* .
	E. n. nordenskioldi Eisen, 1879 *	E. n. caucasica Michaelsen, 1903	E. n. acystis Michaelsen, 1903	E. n. manshurica Kobayashi, 1940	E. n. mongol Blakemore, 2013	E. n. polypapillata Perel, 1969	E. n. onon Blakemore, 2013	E. n. shenzi ssp. nov.	E. n. jilinensis ssp. nov.	E. n. chinensis ssp. nov.
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*Features from Kobayashi’s (1940, 1941) descriptions compared to Zicsi’s.
Length (mm)	25−45 (44–120*)	110	-	111−144	80−110	65−70	100	38−81	40−49	30−50
Diameter	-	4	-	-	-	4-5	-	5.32−6.5	3.57−4.11	3.9−5.1
Segment	106–125 (101– 176*)	107	126	154-175	131–142	102–137	170	78-156	98-127	92-127
Color	Dark puce to pale	D: single-colored, dark, viole-red.	unpigmented,	Dark pigment	Pink-grey	Pale	Reddish pink anterior-dorsum to segment xv otherwise unpigmented	Entire dorsal side is dark-purple except the clitellum (xxv,xxvi −xxxiii), which is gray; Ventro-anterior to xv is purple,>xv is pale	Dorso-anterior to xv pinkish,>xv gradually paling V:pale	D:pale V:pale
Clitellum	xxvi,xxvii−xxxiii	xxiv, xxv−xxxiii	xxvi−xxxiii	xxvi, xxvii−xxxiv	½xxv, xxvi−xxxiii	xxvi, xxvii−xxxii	pale from xxiv dorsally or laterally xxv−xxxiii, i.e., xxiv, xxv−xxxiii	xxv,xxvi−xxxiii	½ xxvi,xxvii−xxxiii	½ xxvi,xxvii−xxxii or xxvi,xxvii−xxxiii
TP	xxix−xxxi	xxvii, xxviii−xxxi or xxviii−½xxxii	xxix−½xxxi	xxix−xxxii	xxix−xxxi	½xxviii, xxix−½xxxi,xxxi	xxviii−xxxi	½xxviii,xxix−xxxi	xxviii,xxix−xxxi or xxix−½xxxii	xxviii,xxix−xxxi or ½xxviii,xxix-½xxxii
Papillae	ab viii,xxvii-xxxi	-	-	ab xxii-xxxv	-	ab xvi-xviii,xxi-xxiii,xxvi,xxxii-xxxiv	-	ab vii-xii cd viii-xii	no	ab viii,xi-xiii cd ix-xiii
Gizzard	-	-	-	-	-	-	xvii−xviii	xvii−xviii,xix	xvii−xviii	xvii−xviii
Setal ratio on segment viii	-	-	-	-	-	-	-	aa:ab:bc:cd:dd = 1.95: 0.34: 1.54: 0.27: 6.30	aa:ab:bc:cd:dd = 1.20: 0.19: 0.92: 0.19: 3.98	aa:ab:bc:cd:dd = 1.48: 0.22: 1.13: 0.21: 4.18
Setal ratio on segment xxviii	-	-	-	-	-	-	-	aa:ab:bc:cd:dd = 3.23: 0.33: 1.39: 0.24: 5.50	aa:ab:bc:cd:dd = 1.96: 0.24: 1.29: 0.17: 4.02	aa:ab:bc:cd:dd = 2.33: 0.32: 1.32: 0.20: 4.25
Setal ratio on segment xli	-	-	-	-	-	-	-	aa:ab:bc:cd:dd = 2.46: 0.29: 1.77: 0.25: 5.31	aa:ab:bc:cd:dd = 1.82: 0.26: 1.22: 0.16: 3.97	aa:ab:bc:cd:dd = 2.00: 0.26: 1.47: 0.22: 3.68

TABLE 5. Morphological comparison of the subspecies of Eisenia nordenskioldi .

E. n. nordenskioldi Eisen, 1879 *

E. n. caucasica Michaelsen, 1903

E. n. acystis Michaelsen, 1903

E. n. manshurica Kobayashi, 1940

E. n. mongol Blakemore, 2013

E. n. polypapillata Perel, 1969

E. n. onon Blakemore, 2013

E. n. shenzi ssp. nov.

E. n. jilinensis ssp. nov.

E. n. chinensis ssp. nov.

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*Features from Kobayashi’s (1940, 1941) descriptions compared to Zicsi’s.

Length (mm)

25−45 (44–120*)

110

111−144

80−110

65−70

100

38−81

40−49

30−50

Diameter

4-5

5.32−6.5

3.57−4.11

3.9−5.1

Segment

106–125 (101– 176*)

107

126

154-175

131–142

102–137

170

78-156

98-127

92-127

Color

Dark puce to pale

D: single-colored, dark, viole-red.

unpigmented,

Dark pigment

Pink-grey

Pale

Reddish pink anterior-dorsum to segment xv otherwise unpigmented

Entire dorsal side is dark-purple except the clitellum (xxv,xxvi −xxxiii), which is gray; Ventro-anterior to xv is purple,>xv is pale

Dorso-anterior to xv pinkish,>xv gradually paling V:pale

D:pale V:pale

Clitellum

xxvi,xxvii−xxxiii

xxiv, xxv−xxxiii

xxvi−xxxiii

xxvi, xxvii−xxxiv

½xxv, xxvi−xxxiii

xxvi, xxvii−xxxii

pale from xxiv dorsally or laterally xxv−xxxiii, i.e., xxiv, xxv−xxxiii

xxv,xxvi−xxxiii

½ xxvi,xxvii−xxxiii

½ xxvi,xxvii−xxxii or xxvi,xxvii−xxxiii

xxix−xxxi

xxvii, xxviii−xxxi or xxviii−½xxxii

xxix−½xxxi

xxix−xxxii

xxix−xxxi

½xxviii, xxix−½xxxi,xxxi

xxviii−xxxi

½xxviii,xxix−xxxi

xxviii,xxix−xxxi or xxix−½xxxii

xxviii,xxix−xxxi or ½xxviii,xxix-½xxxii

Papillae

ab viii,xxvii-xxxi

ab xxii-xxxv

ab xvi-xviii,xxi-xxiii,xxvi,xxxii-xxxiv

ab vii-xii cd viii-xii

ab viii,xi-xiii cd ix-xiii

Gizzard

xvii−xviii

xvii−xviii,xix

xvii−xviii

Setal ratio on segment viii

aa:ab:bc:cd:dd = 1.95: 0.34: 1.54: 0.27: 6.30

aa:ab:bc:cd:dd = 1.20: 0.19: 0.92: 0.19: 3.98

aa:ab:bc:cd:dd = 1.48: 0.22: 1.13: 0.21: 4.18

Setal ratio on segment xxviii

aa:ab:bc:cd:dd = 3.23: 0.33: 1.39: 0.24: 5.50

aa:ab:bc:cd:dd = 1.96: 0.24: 1.29: 0.17: 4.02

aa:ab:bc:cd:dd = 2.33: 0.32: 1.32: 0.20: 4.25

Setal ratio on segment xli

aa:ab:bc:cd:dd = 2.46: 0.29: 1.77: 0.25: 5.31

aa:ab:bc:cd:dd = 1.82: 0.26: 1.22: 0.16: 3.97

aa:ab:bc:cd:dd = 2.00: 0.26: 1.47: 0.22: 3.68

persistent identifier: http://table.plazi.org/id/82B0665A9822FFAD8A8EA8E5A12E4FC4

Published as part of Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025). Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244. DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2 on pages 234-235