Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China Author Qin, Mingyan 0000-0003-4243-9671 State Environmental Protection Key Laboratory of Wetland Ecology and Vegetation Restoration, School of Environment, Northeast Normal University, Changchun 130117, China. zhaohf@lfnu.edu.cn Author Zhao, Huifeng 0000-0003-4243-9671 Key Laboratory of Wetland Ecology and Environment, State Key Laboratory of Black Soils Conservation and Utilization, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, China. zhaohf@lfnu.edu.cn Author Zhang, Yufeng 0000-0003-4243-9671 zhaohf@lfnu.edu.cn Author Han, Anne Charis N. 0000-0003-4243-9671 State Environmental Protection Key Laboratory of Wetland Ecology and Vegetation Restoration, School of Environment, Northeast Normal University, Changchun 130117, China. zhaohf@lfnu.edu.cn Author Shekhovtsov, Sergei V. Department of Molecular Biotechnology, Institute of Cytology and Genetics of the Siberian Branch of the Russian Academy of Sciences, Novosibirsk, Russia. Author Wu, Donghui State Environmental Protection Key Laboratory of Wetland Ecology and Vegetation Restoration, School of Environment, Northeast Normal University, Changchun 130117, China. & Key Laboratory of Wetland Ecology and Environment, State Key Laboratory of Black Soils Conservation and Utilization, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, China. Author Aspe, Nonillon M. College of Fisheries and Marine Sciences, Mindanao State University at Naawan, Naawan 9023, Misamis Oriental, Philippines. text Zootaxa 2025 2025-05-29 5642 3 225 244 https://doi.org/10.11646/zootaxa.5642.3.2 journal article 310169 10.11646/zootaxa.5642.3.2 da3b34c2-b7c6-490d-aaac-3de2beeac5ec 1175-5326 15563696 ED12DD13-CE68-431E-ADE0-050EEEE3EF40

TABLE 4. Morphological comparison of the species of Eisenia.

	E. sibirica Perel & Graphodatsky, 1984	E. lagodechiensis Michaelsen, 1910 (HNHM11047)	E. nana Perel, 1985	E. ventripapillata Manter, 1940	E. atlavinitae Perel & Graphodatsky, 1984	E. tracta Perel, 1985	E. angusta Perel, 1994	E. balatonica Pop, 1943	E. fetida Savigny, 1826	E. nordenskioldi Eisen, 1879 *
Diameter	2.5-4	5	2.5-3	3-4	4-6	5-12	2.5-3	1.5-2.5	2-4	3.57-6.5
Colour	-	-	-	-	Dark purple	-	segments xv-xx have a weak purple pigmentation, strongly blurred on the sides of the ix-xi segments	Sienna-brown	dark violet sometimes alternating stripes of darker segmental with paler intersegmental areas	Unpigmented to pale to different shades of purple
1st dorsal pore	-	-	-	-	4/5	-	4/5	4/5	3/4, 5/6	4/5
Clitellum	xxvii-xxxii, 1/2xxxiii	xxv-xxxiv	xxvii, 1⁄2 xxvii-xxxiii	xxvi-xxxiii	xxvi(xxvii)- xxxiii	xxvi-xxxiii	xxvii-1⁄2 xxxiii	xxiv, xxv–xxx	(xxiv), xxv, xxvi, xxvii–xxxi, xxxii, (xxxiii)	xxiv-xxxiv
Tubercles	xxix-xxx	1⁄2xxviii-1⁄2xxxii	xxix-xxxii	1⁄2xxviii,xxviii-xxx,1⁄2xxxi	xxviii, 1⁄2xxviii-xxxi	xxix-1⁄2xxxii	1⁄2xxix-1⁄2xxxi	xxvi-xxix	1/2 xxviii, xxviii–xxx, xxxi	xxvii-xxxii
Papillae	ab viii, xxviii-xxxi, cd ix-xi	ab xxv-xxxviii, cd ix-xi	ab viii-xii, cd x, xi	ab x, xi, xxvi-xxxii cd x, (xi)	ab vii, viii, xi-xiii, xxvi-xxxiii, xxxiv cd ix-xi	ab viii-x, xvi, xvii, xxiii-xxxvi, xxxvii cd xi, xii	ab xi, xii, xvi, xxvi-xxxii	ab x, xi, xxii, xxiii	abcd ix, x, xi	ab vii-xiii, xvixviii, xxi-xxxv cd viii-xiii
Calciferous glands	-	-	-	-	xi-xii no diverticula	-	xi-xii large no diverticula	xi, xii with diverticula	x-xii no diverticula	xi-xii no diverticula