A review of the flying fish genus Cypselurus (Beloniformes: Exocoetidae). Part 2. Revision of the subgenus Poecilocypselurus Bruun, 1935 with descriptions of three new species and five new subspecies and reinstatement of Exocoetus apus Valenciennes and E. neglectus Bleeker Author Shakhovskoy, Ilia B. Shirshov Institute of Oceanology, Russian Academy of Sciences (IORAS) Nahimovskiy Prospect 36, Moscow, Russia, 117997 ilisha@yandex.com Author Parin, Nikolay V. Shirshov Institute of Oceanology, Russian Academy of Sciences (IORAS) Nahimovskiy Prospect 36, Moscow, Russia, 117997 text Zootaxa 2022 2022-03-23 5117 1 1 109 http://dx.doi.org/10.11646/zootaxa.5117.1.1 journal article 10.11646/zootaxa.5117.1.1 4d31b234-af7e-403f-bc87-f7101f082487 1175-5326 6378619 CFA7895B-43A7-4E19-8623-E8EAE4C43A89
TABLE 1. Dorsal, anal and pectoral-fin ray counts for Cypselurus starksi, C. oligolepis, C. izumii sp. nov., C. clariangulatus sp. nov., C. neglectus, C. nossibe sp. nov. and Cypselurus sp. (poecilopterus?).
Species (subspecies), localityDAPSpecies (subspecies), localityStrD-ASpecies (subspecies), localitySpredSpecies (subspecies), localitySp.bre.brh.brSpecies (subspecies), localityVertVert.pcVert.c
910111213147891013141516176.577.588.599.51010.52345621222324252627282930313233192021222324252645678141516171819383940414243442425262728291314151617
Cypselurus starksi---41631148--13151Cypselurus starksi---13582--18122Cypselurus starksi-----12862121Cypselurus starksi-1166--1-4831-176-1Cypselurus starksi----1104---3102-195-
C. o. oligolepis (South China Sea)--12511--1918--7184-C. o. oligolepis (South China Sea)---38-2--111243-C. o. oligolepis (South China Sea)-12581471-----C. o. oligolepis (South China Sea)-16852---9103--31081-C. o. oligolepis (South China Sea)--232---141---141-
C. o. oligolepis (Singapore)--22010-11913--51971C. o. oligolepis (Singapore)-1--1954---101741C. o. oligolepis (Singapore)-1251293------C. o. oligolepis (Singapore)--711831--7203--216113-C. o. oligolepis (Singapore)--361---271---19--
C. o. oligolepis (Indonesia and Australia)--22313--1424-13149-C. o. oligolepis (Indonesia and Australia)-1171621--113222-C. o. oligolepis (Indonesia and Australia)-12141333------C. o. oligolepis (Indonesia and Australia)-38781--21411---47152-C. o. oligolepis (Indonesia and Australia)--2941--2113---1114-
C. o. oligolepis (Philippines)-110295-12816--5186-C. o. oligolepis (Philippines)---1621--121221-C. o. oligolepis (Philippines)-2-61072------C. o. oligolepis (Philippines)-1-377-1--13511-666-C. o. oligolepis (Philippines)--8163---10102---2155-
C. o. oligolepis (Andaman Sea and Padang Hr.)--3131-1133--381-C. o. oligolepis (Andaman Sea and Padang Hr.)---112----1213--C. o. oligolepis (Andaman Sea and Padang Hr.)--11553------C. o. oligolepis (Andaman Sea and Padang Hr.)1122121--443-13322-C. o. oligolepis (Andaman Sea and Padang Hr.)---92----83---110--
C. o. oligolepis (Bay of Bengal and Arabian Sea)--186--78--26--C. o. oligolepis (Bay of Bengal and Arabian Sea)---1511---3101-C. o. oligolepis (Bay of Bengal and Arabian Sea)---16421-----C. o. oligolepis (Bay of Bengal and Arabian Sea)111-31--1132-1133--C. o. oligolepis (Bay of Bengal and Arabian Sea)--224---411----411
C. o. oligolepis (all regions)-12011946-310282-12585271C. o. oligolepis (all regions)-211366109--460108111C. o. oligolepis (all regions)-57325442202-----C. o. oligolepis (all regions)27243132162133561161313454514-C. o. oligolepis (all regions)-11845161--194111---653111
C. o. persicus---54---81--35-C. o. persicus----11312-63--C. o. persicus-----12122---C. o. persicus--13111--331---232-C. o. persicus----21---2-1---12-
C. o. georgii--96--4101---410-C. o. georgii----131----852-C. o. georgii---2112-------C. o. georgii-122323--552--13442C. o. georgii-273---254----452-
C. o. apus (Philippines)--38---92-1-2--C. o. apus (Philippines)--112----137--C. o. apus (Philippines)-22112-------C. o. apus (Philippines)---1-2----3----1-2-C. o. apus (Philippines)--9----18------81-
C. o. apus (Indonesia)--98--386--4102-C. o. apus (Indonesia)25172----3761-C. o. apus (Indonesia)11276--------C. o. apus (Indonesia)-1-142--1-7----242-C. o. apus (Indonesia)1323---233---125--
C. o. apus (New Guinea)--59--194--372-C. o. apus (New Guinea)---35----2561-C. o. apus (New Guinea)--12351------C. o. apus (New Guinea)--1-71-1-244---3511C. o. apus (New Guinea)--2------2----2---
C. o. apus (Melanesia)--1373-3164--3163-C. o. apus (Melanesia)--2811-----1463-C. o. apus (Melanesia)-1286312-----C. o. apus (Melanesia)-152832--41421-36103-C. o. apus (Melanesia)-292----85----742-
C. o. apus (all regions)--30323-74216-110357-C. o. apus (all regions)2541920----629255-C. o. apus (all regions)14718161022-----C. o. apus (all regions)-26419821162861-3121981C. o. apus (all regions)15225---31910---111173-
C ypselurus oligolepis (species total)-15916253-141541071235127491Cypselurus oligolepis (species total)2753210012121210103141181Cypselurus oligolepis (species total)191452815524522---Cypselurus oligolepis (species total)2113340552782549972523176271283Cypselurus oligolepis (species total)184753182-54357111-12176181
Cypselurus izumii--1551-372--444Cypselurus izumii---17211-1461-Cypselurus izumii----2522-1---Cypselurus izumii---641----92---74--Cypselurus izumii----481--274---58-
Cypselurus clariangulatus--2017--3249--318131Cypselurus clariangulatus--141741--51516--Cypselurus clariangulatus--188131------Cypselurus clariangulatus---421110---13122--510122Cypselurus clariangulatus-116111---792---612--
C. n. neglectus (Tonkin Bay)--1511---169--1158-C. n. neglectus (Tonkin Bay)---21712--6127--C. n. neglectus (Tonkin Bay)----25665----C. n. neglectus (Tonkin Bay)--1446---2481-11110--C. n. neglectus (Tonkin Bay)---82----262--82--
C. n. neglectus (other areas)-125151-4317--6284-C. n. neglectus (other areas)---72032--422151-C. n. neglectus (other areas)--21613154-----C. n. neglectus (other areas)--661173--52071-2121541C. n. neglectus (other areas)--1124---2871--7811
C. n. neglectus (all regions)-140261-44716--74312-C. n. neglectus (all regions)---93744--1034221-C. n. neglectus (all regions)--2181821105----C. n. neglectus (all regions)--71015133--724152-3232541C. n. neglectus (all regions)--1206---210133--151011
C. n. shcherbachevi (Indonesia)--7---331--132-C. n. shcherbachevi (Indonesia)----4----133--C. n. shcherbachevi (Indonesia)----23-1-----C. n. shcherbachevi (Indonesia)--1-32----33--1221-C. n. shcherbachevi (Indonesia)---2------2---2---
C. n. shcherbachevi (Andaman Sea)--1511--3202--3192-C. n. shcherbachevi (Andaman Sea)--1119-----14101-C. n. shcherbachevi (Andaman Sea)----25106-----C. n. shcherbachevi (Andaman Sea)-216731--3116-136101-C. n. shcherbachevi (Andaman Sea)-1344----441-153--
C. n. shcherbachevi (Bay of Bengal and Arabian Sea)--1617--9154--1187-C. n. shcherbachevi (Bay of Bengal and Arabian Sea)---2211---211105-C. n. shcherbachevi (Bay of Bengal and Arabian Sea)-----713102----C. n. shcherbachevi (Bay of Bengal and Arabian Sea)-134872---11131148102-C. n. shcherbachevi (Bay of Bengal and Arabian Sea)--1113----29---83--
C. n. shcherbachevi (all regions)--3828--15387--54011-C. n. shcherbachevi (all regions)--13441---328236-C. n. shcherbachevi (all regions)----41513172----C. n. shcherbachevi (all regions)-351018123--3252212816224-C. n. shcherbachevi (all regions)-14177----6151-1156--
Cypselurus neglectus (species total)-178541-198523--128323-Cypselurus neglectus (species total)--1128154--1362457-Cypselurus neglectus (species total)--21123334277----Cypselurus neglectus (species total)-3122033256--1049373211394781Cypselurus neglectus (species total)-153713---216284-13016--
Cypselurus nossibe--1101--111---471Cypselurus nossibe---173---1-101-Cypselurus nossibe-----23212---Cypselurus nossibe1-321----151-1521--Cypselurus nossibe---144----2411231-
Cypselurus sp. (poecilopterus?)--52514-1358---10277Cypselurus sp. (poecilopterus?)----19971--102211-Cypselurus sp. (poecilopterus?)---313167------Cypselurus sp. (poecilopterus?)--1891272--13188-216138-Cypselurus sp. (poecilopterus?)-21213---116----43-1