An inordinate fondness for inconspicuous brown frogs: integration of phylogenomics, archival DNA analysis, morphology, and bioacoustics yields 24 new taxa in the subgenus Brygoomantis (genus Mantidactylus) from Madagascar Author Scherz, Mark D. Author Crottini, Angelica CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal Author Hutter, Carl R. 0000-0001-6381-6339 Museum of Natural Sciences and Department of Biological Sciences, Louisiana State University, Baton Rouge, LA 70803, USA Ễcarl. hutter @ gmail. com; https: // orcid. org / 0000 - 0001 - 6381 - 6339 USAỄcarl.hutter@gmail.com Author Hildenbrand, Andrea Hauptstr. 13, 82234 Wessling, Germany Ễandrea @ gutachten-hildenbrand. de GermanyỄandrea@gutachten-hildenbrand.de Author Andreone, Franco Museo Regionale di Scienze Naturali, Via G. Giolitti, 36, 10123 Torino, Italy Author Fulgence, Thio Rosin 0000-0001-7205-7282 Natural and Environmental Sciences, Regional University Centre of the SAVA Region (CURSA), Antalaha, Madagascar Ễthiorosinf @ yahoo. fr; https: // orcid. org / 0000 - 0001 - 7205 - 7282 MadagascarỄthiorosinf@yahoo.fr Author Köhler, Gunther Author Ndriantsoa, Serge Herilala Author Ohler, Annemarie Author Preick, Michaela Author Rakotoarison, Andolalao Mention Zoologie et Biodiversité Animale, Université d’Antananarivo, BP 906, Antananarivo, 101 Madagascar Author Rancilhac, Loïs Author Raselimanana, Achille P. Mention Zoologie et Biodiversité Animale, Université d’Antananarivo, BP 906, Antananarivo, 101 Madagascar Author Riemann, Jana C. Author Rödel, Mark-Oliver Author Rosa, Gonçalo M. Author Streicher, Jeffrey W. Author Vieites, David R. Author Köhler, Jörn Author Hofreiter, Michael Author Glaw, Frank Author Vences, Miguel text Megataxa 2022 2022-12-15 7 2 113 311 http://dx.doi.org/10.5281/zenodo.6687412 journal article 217510 10.11646/megataxa.7.2.1 8af27bde-abaa-4f0a-8f84-8b6cc8a97034 2703-3090 7441023 2FD8C310-6486-4592-92F6-5EB894EBD6AC

TABLE 6. Morphometric measurements (all in mm) of voucher specimens of the Mantidactylus ulcerosus clade. Type status is given in square brackets after voucher number: HT, holotype; PT, paratype; LT, lectotype. An asterisk (*) marks lectotypes designated in the current paper. A hash (#) marks measurements taken by AH and thus not fully comparable with other measurements, all taken by MV. For abbreviations of measurements, see Materials and Methods. NM, not measured; NA, not applicable.

Voucher	Field number	Sex	Locality	SVL	HW	HL	ED	HTD	END	NSD	NND	FORL	HAL	HIL	FOTL	FOL	TIBL	FGL	FGW
M. ulcerosus
...Continued on the next page
...Continued on the next page
...Continued on the next page
...Continued on the next page
...Continued on the next page
...Continued on the next page
...Continued on the next page
...Continued on the next page
...Continued on the next page
...Continued on the next page
...Continued on the next page
SMF 6605 [LT*]	NA	F	Nosy Be	45.4	17.8	18.7	6.9	4.9	4.4	2.5	4.2	26.2	12.1	65.8	30.3	21.7	20.6	NA	NA
ZMB 53737 [LT brauni *]	NA	M	Uncertain (Akkoraka)	28.5	11.4	11.7	3.9	3.1	3.7	1.4	2.5	17.9	8.2	NM	NM	NM	14.8	NM	NM
ZFMK 52659 #	NA	F	Nosy Be	32.5	12.5	11.3	4.7	3.7	3.2	2.2	3.7	11.5	9.4	NM	NM	16.0	15.9	2.5	2.2
ZFMK 52666 #	NA	F	Nosy Be	38.7	15.7	13.2	5.3	2.8	4.1	1.9	4.0	13.5	11.5	NM	NM	17.8	19.2	NA	NA
ZFMK 52667 #	NA	F	Nosy Be	40.7	17.4	14.8	5.7	3.2	4.7	2.8	4.3	16.2	12.1	NM	NM	21.5	18.8	2.1	1.5
ZFMK 52668 #	NA	F	Nosy Be	40.6	16.3	14.6	5.6	3.7	4.1	2.7	4.3	11.6	11.1	NM	NM	19.3	18.7	NA	NA
ZFMK 53668 #	NA	F	Nosy Be	40.3	15.8	14.7	5.4	3.6	4.2	2.8	4.4	15.0	12.1	NM	NM	19.3	19.4	1.8	1.4
ZSM 407/2000 #	FGMV 2000.437	F	Benavony	37.4	15.6	15.7	5.4	3.9	3.7	2.5	4.6	14.4	12.5	NM	NM	18.7	19.1	1.3	1.1
ZSM 408/2000 #	FGMV 2000.438	F	Benavony	40.0	16.6	15.6	5.4	3.7	4.5	2.2	4.4	15.4	12.2	NM	NM	19.5	20.2	1.1	1.1
ZSM 563/2009	ZCMV 11465	F	Makira (Sa- haovy)	42.1	16.0	17.7	5.6	4.4	4.1	2.2	4.5	22.4	11.5	61.8	27.5	19.9	16.9	NA	NA
ZSM 566/2009	ZCMV 11481	F	Makira (Sa- haovy)	40.5	15.9	16.7	5.3	3.8	3.9	2.8	4.0	24.2	12.3	61.5	27.9	19.9	18.7	NA	NA
ZSM 568/2009	ZCMV 11459	F	Makira (Sa- haovy)	43.3	17.5	17.3	5.8	3.1	4.2	2.5	4.4	24.3	13.2	61.0	27.7	19.5	18.1	NA	NA
ZSM 599/2001 #	FGMV 2001.20	F	Benavony	40.3	16.4	15.1	6.0	3.4	4.6	2.5	3.9	13.9	12.8	NM	NM	19.3	19.5	NA	NA
ZSM 707/2001	FGMV 2001.270	F	Ankarafant- sika	40.7	14.9	16.9	5.6	3.9	4.2	2.8	3.4	23.1	10.9	57.3	25.5	17.4	17.6	NA	NA
ZSM 989/2001 #	NA	F	Ankarafant- sika	34.7	13.6	12.0	4.3	3.0	3.6	2.3	3.7	12.1	8.5	NM	NM	15.2	16.6	1.3	1.1
ZFMK 53669 #	NA	M	Nosy Be	32.2	13.7	12.4	4.4	4.1	3.5	2.0	3.5	10.7	9.2	NM	NM	14.2	16.5	3.4	2.5
ZFMK 53670 #	NA	M	Nosy Be	33.8	13.6	12.5	4.6	4.0	3.8	2.2	4.0	12.0	9.3	NM	NM	14.4	15.5	3.5	2.9
ZSM 565/2009	ZCMV 11478	M	Makira (Sa- haovy)	34.6	13.3	14.0	5.0	3.7	3.4	2.6	3.4	19.4	9.5	51.2	23.0	16.6	14.6	5.7	3.0
ZSM 590/2001	FGMV 2001.4	M	Nosy Be	35.9	13.8	15.6	5.5	5.2	3.8	2.3	3.6	19.5	10.6	51.0	22.9	16.5	15.9	7.0	4.4
ZSM 598/2001 #	FGMV 2001.19	M	Benavony	32.9	13.9	13.0	5.2	4.5	3.8	2.1	3.3	12.7	10.2	NM	NM	14.8	15.1	3.1	2.1
ZSM 708/2001	FGMV 2001.400	M	Ankarafant- sika	33.9	14.3	14.7	5.1	4.4	3.9	2.7	3.7	19.4	9.5	50.1	22.3	16.2	14.2	6.7	2.9
ZSM 709/2001 #	FGMV 2001.401	M	Ankarafant- sika (Ampijoroa)	31.7	13.7	12.5	4.3	4.1	3.6	2.1	3.2	12.1	9.5	NM	NM	11.2	14.6	4.2	2.6
M. bellyi
MNHN 1983.240 [HT]	NA	F	Montagne d’Ambre	36.7	15.4	15.9	5.2	3.9	NM	NM	NM	21.2	11.5	56.0	26.4	18.2	16.3	1.3	1.3
ZFMK 52660 #	NA	F	Andrakata	39.1	16.9	13.9	5.7	3.8	3.9	2.6	4.1	13.8	8.9	NM	NM	16.1	18.3	4.0	3.3
ZFMK 52661 #	NA	F	Andrakata	40.6	15.5	13.6	5.3	2.8	3.4	2.5	4.1	13.5	11.5	NM	NM	18.9	21.0	NA	NA
ZFMK H14140 #	NA	F	‘Diego-is- land’ (near Antsiranana)	37.9	15.1	13.3	5.5	3.1	3.5	2.0	2.8	14.0	10.6	NM	NM	17.0	18.0	NA	NA
ZFMK H14142 #	NA	F	‘Diego-is- land’ (near Antsiranana)	37.0	15.4	13.9	4.8	3.7	4.3	1.6	3.9	NM	11.7	NM	NM	17.5	19.4	NA	NA
ZFMK H14143 #	NA	F	‘Diego-is- land’ (near Antsiranana)	33.0	13.5	12.6	4.9	3.6	3.8	1.9	3.8	10.1	9.4	NM	NM	15.6	17.1	NA	NA
ZSM 197/2004 #	FGZC 370	F	Montagne d’Ambre ANGAP House	46.4	18.0	16.9	5.9	4.1	4.3	3.0	4.7	17.0	12.6	NM	NM	20.1	22.2	NA	NA
ZSM 292/2004 #	FGZC 560	F	Montagne des Francais, Andavakoera	41.3	19.0	14.2	5.1	3.8	3.9	2.5	3.7	14.1	12.3	NM	NM	20.1	20.5	NA	NA
ZSM 293/2004 #	FGZC 563	F	Montagne des Francais, Andavakoera	44.2	19.0	15.9	5.4	4.1	4.5	2.4	3.8	14.1	12.6	NM	NM	18.8	19.4	2.2	1.6
ZSM 358/2004 #	FGZC 411	F	Montagne dÀmbre	44.4	18.9	15.9	5.3	3.8	4.2	2.9	4.2	15.1	13.2	NM	NM	20.4	20.3	NA	NA
ZSM 755/2003 #	FGMV 2002.549	F	Ankarana	43.6	18.7	15.9	5.7	3.4	4.7	2.8	4.5	13.9	12.6	NM	NM	18.6	19.7	NA	NA
ZSM 756/2003 #	FGMV 2002.551	F	Ankarana	43.3	16.9	16.5	5.6	4.2	4.2	2.4	4.4	14.8	13.1	NM	NM	19.6	19.9	NA	NA
ZFMK 62201 #	NA	M	Ankarana	33.3	15.1	13.0	4.9	5.0	3.4	2.5	4.1	12.0	9.1	NM	NM	14.4	15.4	2.8	2.2
ZFMK H14141 #	NA	M	‘Diego-is- land’ (near Antsiranana)	31.7	14.8	12.0	5.0	4.5	3.5	2.4	3.7	10.8	10.6	NM	NM	14.0	16.2	4.4	2.4
ZSM 208/2004	FGZC 408	M	Montagne d’Ambre	40.1	15.8	15.9	5.6	5.5	3.0	2.0	3.8	21.7	11.4	57.3	25.4	18.3	17.0	3.4	2.4
ZSM 209/2004	FGZC 409	M	Montagne d’Ambre	40.5	16.8	16.1	4.7	5.0	3.9	2.5	4.2	23.0	11.4	59.3	27.0	19.2	17.8	3.9	2.7
ZSM 231/2004 #	FGZC 452	M	Montagne d’Ambre	40.7	20.0	15.4	5.2	5.1	4.5	2.7	3.8	14.6	12.4	NM	NM	18.5	20.1	3.5	2.5
ZSM 291/2004 #	FGZC 559	M	Montagne des Francais, Andavakoera	35.5	16.4	13.3	4.8	4.1	3.7	2.1	3.4	12.1	11.3	NM	NM	16.7	16.7	3.1	2.1
ZSM 294/2004 #	FGZC 564	M	Montagne des Francais, Andavakoera	33.1	15.5	12.5	4.2	4.3	3.4	2.0	3.2	12.1	9.9	NM	NM	15.3	16.2	3.0	2.3
ZSM 295/2004 #	FGZC 566	M	Montagne des Francais, Andavakoera	33.1	14.8	13.0	5.0	4.8	3.4	2.0	2.9	12.9	9.6	NM	NM	15.8	16.1	4.3	2.9
ZSM 323/2005 #	FGZC 2715	M	Andapa	35.2	14.9	13.1	5.1	3.6	3.6	2.4	3.7	12.9	10.6	NM	NM	17.2	18.3	3.0	2.4
ZSM 753/2003	FGMV 2002.545	M	Ankarana	38.1	16.0	16.0	5.6	5.9	3.9	1.9	3.9	19.7	10.7	55.0	25.0	17.8	16.9	4.4	3.3
ZSM 754/2003	FGMV 2002.596	M	Ankarana	41.0	17.6	17.9	6.0	6.8	3.3	2.6	4.2	22.7	11.6	59.8	26.9	NM	17.5	4.6	3.3
Mantidactylus schulzi
ZSM 653/2001 [HT]	MV 2001.150 (HT)	M	Tsaratanàna, Manarikoba	23.5	9.0	10.2	3.4	3.2	2.9	1.6	3.1	19.3	6.6	35.9	15.5	10.5	10.9	5.2	2.7
ZSM 821/2003 [PT]	FGMV 2002.751	M	Manongarivo	22.9	8.7	10.0	3.5	3.2	2.1	1.5	3.0	14.0	6.4	34.9	14.8	10.5	10.8	5.0	2.6
ZSM 822/2003 [PT]	FGMV 2002.752	M	Manongarivo	21.4	8.5	9.5	3.4	3.2	2.7	1.4	3.1	13.9	6.8	34.8	15.2	10.5	10.2	4.9	2.4
ZSM 651/2001 [PT]	MV 2001.136	F	Tsaratanàna, Manarikoba	28.7	10.5	11.6	4.1	3.0	2.7	1.8	3.5	17.9	8.6	44.8	20.4	13.8	14.0	NA	NA
ZSM 652/2001 [PT]	MV 2001.137	F	Tsaratanàna, Manarikoba	26.2	9.5	10.7	3.6	2.7	2.6	1.8	3.0	16.3	8.2	41.1	18.4	12.2	12.7	NA	NA
ZSM 654/2001 [PT]	MV 2001.15	F	Tsaratanàna, Manarikoba	27.3	9.5	10.5	3.8	2.7	2.3	1.6	3.3	15.7	7.1	38.9	16.8	11.3	11.9	NA	NA
ZSM 823/2003 [PT]	FGMV 2002.753	F	Manongarivo	28.9	10.3	11.9	3.9	3.2	2.8	2.0	3.1	17.4	8.2	43.6	19.4	13.4	13.1	NA	NA
ZSM 824/2003 [PT]	FGMV 2002.755	F	Manongarivo	27.1	10.1	10.5	4.0	3.2	2.6	1.7	3.2	16.3	7.6	42.1	18.4	12.3	13.3	NA	NA
ZSM 825/2003 [PT]	FGMV 2002.757	F	Manongarivo	24.7	9.6	10.8	3.5	2.5	2.4	1.8	3.3	15.5	7.4	39.6	17.6	11.9	12.7	NA	NA
ZSM 826/2003 [PT]	FGMV 2002.759	F	Manongarivo	28.5	10.3	11.0	3.5	2.8	2.8	1.8	3.4	15.2	7.6	39.7	18.5	12.5	12.6	NA	NA
Mantidactylus stein- fartzi sp. nov. (Ca33)
ZSM 658/2001 [HT]	FGMV 2001.107	M	Manarikoba	21.5	8.9	9.2	3.3	3.5	1.8	1.9	2.6	13.9	6.4	30.9	13.6	9.3	9.6	3.9	2.6
M. betsileanus
BMNH 1947.2.26.45 [LT*]	NA	M	Betsileo and Ankafana	22.8	7.6	8.9	3.2	2.3	2.4	1.5	2.5	13.5	6.0	40.1	17.8	12.8	NM	2	1.6
BMNH 1947.2.26.33 # [PLT]	NA	F	Betsileo and Ankafana	31.5	11.8	10.5	4.4	2.5	2.9	2.1	3.3	11.3	9.3	NM	NM	17.5	18.8	NA	NA
BMNH 1947.2.26.34 # [PLT]	NA	F	Betsileo and Ankafana	30.3	11.5	10.4	4.2	3.0	3.0	2.3	3.4	11.5	9.5	NM	NM	16.3	17.5	1.1	1.0
BMNH 1947.2.26.41 # [PLT]	NA	F	Betsileo and Ankafana	30.7	11.9	10.5	4.4	2.9	2.9	1.9	3.5	10.3	8.6	NM	NM	14.9	16.9	0.8	0.7
BMNH 1947.2.26.42 # [PLT]	NA	F	Betsileo and Ankafana	29.5	11.1	10.4	3.8	2.7	3.2	2.2	3.3	10.2	8.8	NM	NM	16.5	17.7	1.3	0.9
BMNH 1947.2.26.44 [PLT]	NA	F	Betsileo and Ankafana	30.0	10.2	10.9	4.0	2.3	2.8	2.0	3.3	17.5	8.6	57.4	26.6	17.7	NM	NA	NA
ZMB 30514 [HT brunneus]	NA	F	Probably eastern or central Madagascar	31.4	10.8	11.8	4.0	2.7	2.9	1.9	3.5	17.0	8.2	53.5	24.5	16.2	16.6	NA	NA
SMF 6733 [HT multiplicatus]	NA	F	Lac Alaotra	36.2	12.8	14.6	4.6	3.0	3.8	2.4	3.6	19.5	9.5	59.9	27.9	19.6	19.9	NA	NA
MNHN 1895.258 [HT fumigatus]	NA	F	West coast of Madagascar	33.2	11.8	13.6	3.5	2.4	3.2	2.5	3.1	NM	NM	NM	NM	NM	NM	NA	NA
MRSN A7042	FAZC 15565	F	Maromizaha	35.5	12.4	13.6	4.6	3.6	3.0	2.4	3.4	21.4	9.4	62.0	28.5	19.3	18.9	NA	NA
ZSM 200/2021	FAZC 15578	F	Maromizaha	34.4	11.3	14.0	4.7	3.4	2.9	2.5	4.0	20.0	8.9	59.4	21.3	17.4	17.8	NA	NA
ZMA 20206	ZCMV 53	M	Ranomafana	24.3	9.8	8.9	3.5	3.3	2.6	2.0	2.6	8.7	6.6	NM	NM	12.2	13.3	3.0	2.1
ZMA 20210	ZCMV 141	M	Ranomafana	24.1	9.2	8.1	3.3	3.7	2.1	1.6	2.6	7.7	6.1	NM	NM	11.4	12.2	2.7	2.0
ZSM 1772/2008	ZCMV 8017	M	Ranomafana	21.9	8.2	9.6	3.4	3.0	2.0	1.8	2.8	13.9	7.2	39.9	17.7	12.5	12.2	2.5	1.9
ZSM 1773/2008	ZCMV 8068	M	Ambodisakoa	25.0	9.0	10.5	3.8	4.1	2.9	2.1	3.3	15.7	7.4	41.1	19.0	13.3	12.8	3.0	2.0
ZSM 1774/2008	ZCMV 8776	M	Mantadia (Camp Prolemur)	28.5	11.1	12.0	4.0	4.6	2.9	1.8	3.0	16.4	7.8	47.5	22.4	15.2	14.1	3.0	1.5
ZSM 1775/2008	ZCMV 8778	M	Mantadia (Camp Prolemur)	29.0	10.7	12.6	4.5	4.6	2.8	2.0	3.0	15.5	7.6	46.5	21.7	15.2	13.5	2.7	2.3
ZSM 391/2006	ZCMV 3260	M	Andasibe	26.2	10.1	11.7	4.4	4.3	3.3	2.2	3.0	17.1	8.7	51.9	23.8	16.5	15.5	3.0	1.8
ZSM 738/2009	ZCMV 7164	M	Befanjana	28.9	10.4	11.8	4.5	3.8	3.3	1.8	3.0	18.3	8.6	48.2	22.0	14.1	15.6	3.2	2.0
ZSM 739/2009	ZCMV 7293	M	Befanjana	27.4	10.5	12.9	4.4	3.4	2.7	1.9	2.8	17.0	7.9	48.5	22.2	15.0	14.9	3.7	2.3
ZSM 82/2002	FGMV 2001.1049	M	Andasibe	27.8	10.5	10.2	3.8	3.9	2.6	1.8	3.1	10.0	8.0	NM	NM	14.3	14.6	2.6	1.7
M. noralottae
MRSN A5317 * [HT]	NA	M	Isalo	34.8	13.5	15.0	4.9	4.0	3.3	2.5	3.7	17.8	10.0	NM	25.9	18.7	NM	2.2	2.5
MRSN A5036 * [PT]	NA	M	Isalo	35.5	12.9	14.8	5.2	4.2	3.0	2.3	3.7	19.8	11.5	NM	29.8	19.3	NM	2.0	1.7
MRSN A5319 * [PT]	NA	M	Isalo	33.4	12.2	13.0	4.0	3.8	3.2	1.8	3.6	18.3	11.0	NM	28.3	18.3	NM	2.5	2.0
SMF 85861 * [PT]	NA	M	Isalo	32.8	13.5	12.8	5.1	3.2	3.4	2.3	3.7	14.9	9.1	NM	28.6	18.3	NM	2.0	2.0
MRSN A5252 * [PT]	NA	F	Isalo	38.8	14.5	16.6	5.5	3.4	4.4	2.2	4.5	18.8	11.0	NM	27.8	19.9	NM	NA	NA
MRSN A5254 * [PT]	NA	F	Isalo	37.7	13.3	14.5	5.5	3.4	3.4	3.3	4.4	18.8	11.1	NM	26.7	19.9	NM	NA	NA
MRSN A5318 * [PT]	NA	F	Isalo	38.5	15.2	16.3	5.6	3.6	3.2	2.8	4.3	19.8	11.5	NM	30.5	20.5	NM	NA	NA
SMF 85862 * [PT]	NA	F	Isalo	37.8	14.1	14.7	5.9	3.3	3.2	2.8	3.9	17.5	9.6	NM	29.1	18.8	NM	NA	NA
SMF 85863 * [PT]	NA	F	Isalo	35.5	13.3	14.5	4.4	3.2	3.3	3.3	4.4	18.8	11.1	NM	27.0	19.6	NM	NA	NA
SMF 85864 * [PT]	NA	F	Isalo	40.0	15.5	15.5	5.6	3.4	4.4	2.2	3.4	20.0	11.1	NM	31.1	21.1	NM	NA	NA
ZSM 49/2011 [PT]	FAZC 13020 / MRSN A5035	F	Isalo	36.9	13.6	15.2	5.3	3.8	3.7	2.6	4.1	23.1	11.5	62.6	28.4	18.7	19.7	NA	NA
ZSM 188/2021	ACZCV 295	J?	Isalo, Canyon des rats	26.2	9.6	10.9	3.8	2.8	2.5	2.0	3.0	16.2	9.0	44.7	19.6	12.9	13.7	NA	NA
M. tripunctatus (Ca29)
MNHN 1931 23 # [PLT]	NA	J?	St.Louis (Fort Dauphin), Befotaka (Farafangana)	21.1	8.7	8.5	3.3	2.7	2.5	1.3	2.6	6.0	6.1	NM	NM	10.4	11.9	NA	NA
MNHN 1931.24 [LT*]	NA	J?	St.Louis (Fort Dauphin), Befotaka (Farafangana)	23.0	8.5	10.2	3.4	2.0	2.7	1.7	2.6	18.4	6.6	37.2	17.2	12.5	12.0	NA	NA
MNHN 1931.21 [PLT]	NA	F?	St.Louis (Fort Dauphin), Befotaka (Farafangana)	28.7	10.8	12.2	3.2	2.4	3.0	2.5	3.2	17.5	8.3	NM	22.6	15.8	15.6	NA	NA
ZSM 71/2004	FGZC 119	M	Andohahela	26.4	10.3	11.5	4.3	4.0	3.0	1.9	3.1	17.5	8.0	45.6	20.0	13.5	14.3	3.9	2.0
ZFMK 52665 #	NA	M?	Tolagnaro (Fort Dauphin)	27.4	10.8	10.8	4.4	3.6	3.0	1.9	2.8	9.3	7.4	NM	NM	12.7	13.8	2.1	1.8
ZFMK 52663 #	NA	M	Tolagnaro (Fort Dauphin)	26.6	10.7	9.9	4.0	3.7	2.5	1.6	3.0	9.5	8.6	NM	NM	13.0	14.2	3.4	2.0
ZFMK 53666 #	NA	M	Tolagnaro (Fort Dauphin)	27.0	11.1	10.4	3.9	3.7	3.0	2.3	3.1	9.5	7.4	NM	NM	13.7	13.8	3.8	2.4
ZFMK 52664 #	NA	F	Tolagnaro (Fort Dauphin)	35.1	13.6	12.6	4.9	3.6	3.4	3.0	3.5	12.5	9.3	NM	NM	15.9	19.2	NA	NA
ZFMK 53667 #	NA	F	Tolagnaro (Fort Dauphin)	32.9	12.0	12.3	4.8	3.2	3.6	2.4	3.3	NM	NM	NM	NM	15.3	16.8	NA	NA
ZFMK 56185 #	NA	F	Nahampoana	34.1	12.9	12.7	4.9	2.6	3.2	2.3	3.7	11.7	10.1	NM	NM	16.8	17.9	NA	NA
ZSM 172/2004	FGZC 318	F	Manantantely	33.8	12.7	13.3	5.0	3.3	3.5	1.9	3.7	20.8	9.4	59.0	25.4	18.5	18.0	NA	NA
ZSM 70/2004	FGZC 117	F	Andohahela	34.7	12.6	14.4	4.4	4.1	3.1	2.7	3.9	22.6	10.0	62.5	28.4	19.3	19.3	NA	NA
M. incognitus sp. nov. (Ca34)
ZSM 669/2009 [HT]	ZCMV 8945	F	Mahanoro	26.4	10.6	11.7	4.1	2.5	3.0	2.2	3.0	15.3	8.0	48.8	20.9	14.3	15.0	NA	NA
MRSN A6694 [PT]	RJS 1801 (ACZC 4189)	M	Anivorano Est	27.4	10.6	12.0	4.5	3.8	2.4	2.0	3.8	16.2	7.7	44.9	21.4	14.7	14.4	4.1	2.8
M. jonasi sp. nov.
ZSM 86/2016 [HT]	MSZC 0180	M	Ampotsidy	24.2	8.6	9.7	4.3	4.0	2.5	1.9	2.9	14.4	7.2	39.7	18.9	13.0	11.6	2.5	1.7
ZSM 133/2018 [PT]	MSZC 0714	M	Montagne d’Ambre	24.8	10.5	10.8	3.5	4.4	2.6	1.9	3.2	15.6	7.8	42.9	19.6	13.9	13.1	3.0	1.9
ZSM 134/2018 [PT]	MSZC 0715	M	Montagne d’Ambre	24.9	10.0	10.4	3.6	3.6	2.6	1.7	3.0	15.8	7.5	41.6	19.3	13.3	12.8	3.0	2.1
ZSM 1773/2010 [PT]	ZCMV 12603	M	Bealanana-Antsohihy	23.8	9.3	10.4	3.8	3.5	2.5	1.5	2.5	14.2	7.2	40.7	19.0	12.9	11.9	3.0	2.0
ZSM 1774/2010 [PT]	ZCMV 12604	M	Bealanana-Antsohihy	22.8	8.6	9.9	3.5	3.4	2.0	1.8	2.5	14.0	6.9	39.9	18.0	12.6	11.9	2.7	1.7
ZSM 2220/2007 [PT]	FGZC 1368	M	Montagne d’Ambre	23.1	9.2	9.8	3.5	3.8	2.4	1.8	2.4	14.7	7.0	40.9	18.1	12.7	12.6	3.5	2.0
ZSM 2222/2007 [PT]	FGZC 1372	M	Montagne d’Ambre	24.5	10.0	10.8	3.4	3.5	2.7	1.9	2.9	15.0	6.8	42.4	19.8	13.4	13.0	4.0	2.5
ZSM 551/2009 [PT]	ZCMV 11462	M	Makira West	22.4	8.4	9.6	3.1	3.4	2.2	1.5	2.5	13.6	7.5	34.7	17.6	12.1	11.8	3.4	2.2
ZSM 89/2016 [PT]	MSZC 0194	M	Andranonafindra	24.6	9.6	10.4	4.0	4.3	2.6	1.5	2.6	15.1	7.4	41.2	20.1	13.8	12.7	3.9	2.2
M. katae sp. nov. (Ca28)
ZSM 79/2002 [HT]	FGMV 2001.1179 (2002.G24)	M	Andasibe	24.6	9.9	10.3	4.0	2.9	2.5	1.6	2.6	15.7	7.3	43.7	19.4	13.0	13.4	4.1	2.6
ZSM 637/2003 [PT] #	FGMV 2002.132	M	Ranomafana	23.2	9.6	8.7	3.3	2.7	2.3	1.8	2.7	8.2	7.8	NM	NM	12.1	13.6	3.5	2.4
ZSM 668/2003 [PT] #	FGMV 2002.255	M	Ranomafana	24.5	9.4	9.0	4.1	3.1	2.3	1.8	2.6	8.7	7.7	NM	NM	13.0	14.0	4.3	3.3
ZSM 708/2003 [PT] #	FGMV 2002.348	M	Ranomafana	22.1	9.0	8.5	3.2	2.7	2.0	1.3	2.7	7.5	6.6	NM	NM	11.0	12.6	3.8	2.4
ZSM 81/2002 [PT]	FGMV 2001.1275 (2002.G66)	M	Andasibe	26.1	9.1	10.6	3.4	3.2	2.4	1.9	2.6	14.9	7.1	43.5	20.5	14.1	13.5	4.7	2.8
ZSM 91/2004	FGZC 163	M	Andohahela	27.2	9.9	11.5	4.0	3.7	2.8	1.9	3.4	17.3	8.2	46.0	21.6	NM	13.6	4.5	2.7
ZSM 196/2021 [PT]	FAZC 15504	M	Maromizaha	25.6	9.8	10.6	4.0	3.5	2.5	2.2	2.9	15.5	7.1	41.4	18.8	13.6	12.7	4.3	2.9
ZSM 197/2021 [PT]	FAZC 15507	M	Maromizaha	25.5	9.7	10.3	4.1	3.4	2.1	1.9	3.0	15.0	7.9	45.3	20.1	13.7	13.8	4.0	2.9
ZSM 80/2002 [PT] #	FGMV 2001.1180	F	Andasibe	25.6	10.1	9.5	3.7	2.5	2.9	2.3	3.0	10.0	7.7	NM	NM	13.4	15.5	1.4	1.4
ZSM 83/2002 [PT]	FGMV 2001.1173 (2002.G18)	F	Andasibe	35.9	12.8	13.6	5.0	3.4	3.3	2.2	3.8	20.7	9.5	61.8	27.7	18.8	18.5	1.1	1.1
MRSN A7043 [PT]	FAZC 15523	F	Maromizaha	32.5	12.6	13.8	4.4	2.7	3.0	2.5	3.7	20.6	8.7	59.6	27.0	17.7	18.6	NA	NA
M. kortei sp. nov.
ZSM 205/2005 [HT]	FGZC 2376	M	Andohahela	27.4	11.1	11.8	4.1	3.6	2.8	1.9	3.7	16.7	8.4	43.1	20.2	13.9	12.7	5.8	2.6
ZSM 195/2005 [PT]	FGZC 2480	F	Andohahela	34.6	12.3	13.7	4.4	3.2	3.1	2.4	3.9	19.1	9.5	55.6	25.5	17.5	16.9	NA	NA
ZSM 203/2005 [PT]	FGZC 2377	F	Andohahela	37.1	12.6	13.6	4.0	3.2	3.6	2.0	3.1	20.8	9.8	55.0	24.6	17.1	16.7	NA	NA
ZSM 204/2005 [PT]	FGZC 2375	F	Andohahela	35.0	13.5	14.4	4.7	3.7	3.4	2.7	4.3	21.7	10.7	58.3	27.0	18.6	17.6	1.7	1.3
M. riparius sp. nov.
ZSM 2403/2007 [HT]	ZCMV 5766	M	Isalo	27.3	10.5	11.4	4.4	3.6	2.6	2.2	3.0	16.0	8.1	45.1	20.7	14.3	13.7	5.0	2.1
ZSM 186/2021 [PT]	ACZCV 281	F	Isalo: Andremanero	27.3	10.0	11.2	4.0	2.7	2.7	2.0	3.0	16.8	7.7	43.5	19.4	13.5	13.4	NA	NA
ZSM 187/2021 [PT]	ACZCV 283	F	Isalo: Andremanero	21.5	7.3	9.0	3.3	2.0	1.9	1.4	2.6	12.0	5.7	34.2	14.7	10.0	10.5	NA	NA
M. fergusoni sp. nov. (Ca26)
ZSM 126/2002 [HT]	FGMV 2001.1389	M	Andranofotsy	27.8	10.3	11.6	4.4	4.0	3.0	1.9	2.8	17.5	8.7	48.7	22.5	15.1	14.5	3.6	2.6
ZFMK 59938 #	NA	M	Marojejy, Camp Mantella	26.0	10.6	9.7	3.9	2.6	2.9	1.8	2.9	8.9	8.0	NM	NM	13.4	14.5	2.8	2.2
ZSM 125/2002 [PT]	FGMV 2001.1434	M	Andranofotsy	26.2	10.1	11.2	4.2	3.1	2.4	2.0	3.1	15.8	7.7	41.8	19.1	13.3	13.4	3.9	2.4
ZSM 201/2005 #	FGZC 2746	M	Marojejy, Camp Mantella	24.9	9.9	10.0	4.1	3.3	2.7	1.8	2.7	10.2	7.9	NM	NM	12.3	14.0	3.4	2.3
ZSM 5051/2005 [PT]	ZCMV 2136	M	Nosy Mangabe	29.9	11.4	12.8	4.1	3.6	3.5	2.4	3.3	17.6	8.5	45.6	21.0	14.0	14.8	3.7	2.5
ZFMK 59939 #	NA	F	Marojejy, Camp Mantella	35.7	14.7	13.4	5.4	2.9	3.8	2.1	4.0	12.7	10.4	NM	NM	16.3	19.6	NA	NA
ZSM 124/2002 [PT]	FGMV 2001.1433	F	Andranofotsy	36.1	13.6	14.5	5.6	3.6	3.9	2.3	3.1	22.5	10.2	60.2	28.3	19.4	18.9	NA	NA
ZSM 355/2010 [PT]	FGZC 4277	F	Makira	40.3	15.2	16.3	5.5	3.8	4.1	2.4	3.9	24.6	11.8	66.5	30.0	21.2	21.2	NA	NA
ZSM 5050/2005 [PT]	ZCMV 2125	F	Nosy Mangabe	42.2	15.4	16.7	6.0	4.0	4.2	2.7	3.8	24.4	11.9	69.4	30.6	21.0	21.7	NA	NA
M. georgei sp. nov. (Ca35/Ca36)
ZSM 455/2005 [HT]	ZCMV 806	M	Toamasina	30.7	11.9	12.4	4.9	4.7	3.2	1.9	3.5	17.4	8.0	46.2	20.6	14.1	13.7	4.0	3.3
ZSM 454/2005	ZCMV 803	M	Maroantsetra	29.0	10.9	12.0	4.6	3.7	3.3	1.8	3.4	17.4	8.6	46.9	20.3	13.9	13.8	4.0	2.4
ZSM 456/2005 [PT]	ZCMV 807	M	Toamasina	28.4	10.5	11.9	4.4	4.2	2.9	1.8	3.5	16.3	8.4	46.3	20.4	13.8	13.5	4.6	3.3
M. jahnarum sp. nov. (Ca27)
ZSM 423/2006 [HT]	ZCMV 3390	M	Nosy Boraha	30.3	11.1	12.6	4.7	3.3	3.1	1.8	3.2	17.2	8.9	46.1	21.1	14.0	14.4	4.8	3.0
ZSM 424/2006 [PT]	ZCMV 3393	M	Nosy Boraha	28.5	10.5	11.8	4.5	3.1	3.1	1.7	3.0	17.5	9.1	40.8	20.0	13.5	14.1	4.6	3.4
ZSM 520/2006 [PT]	ZCMV 3218	F	Nosy Boraha	33.6	12.5	13.8	4.9	3.6	3.0	2.0	3.0	21.5	9.7	57.2	26.3	17.5	17.1	NA	NA
ZSM 521/2006 [PT]	ZCMV 3219	F	Nosy Boraha	30.4	11.1	12.6	4.7	2.9	3.1	2.0	3.2	19.4	9.7	50.7	23.2	15.6	15.4	NA	NA
M. marintsoai sp. nov.
CURSA-A033/2021 [HT] *	THC301	M	Bangoabe, Marojejy	29.0	10.3	12.1	3.7	3.8	3.4	1.9	2.9	16.4	8.1	44.7	20.2	14.0	14.2	NM	NM
CURSA-A036/2021 [PT] *	THC354	F	Andramanolotra	36.7	12.4	14.6	4.7	3.0	3.5	1.9	3.3	20.6	8.7	60.6	24.6	17.7	17.6	NM	NM
CURSA-A035/2021 [PT] *	THC233	F	Antsirabe-Nord (Bemanevika)	36.8	11.7	14.0	4.2	3.0	3.8	2.3	3.6	21.8	10.8	59.8	25.6	18.2	18.5	NM	NM
CURSA-A034/2021 [PT] *	THC248	F	Andramanolotra	34.6	11.4	13.0	4.1	2.8	3.8	1.9	3.1	20.0	10.0	52.9	23.7	16.4	16.4	NM	NM
ZSM uncatalogued [PT] *	THC246	F	Andramanolotra	36.5	12.7	14.6	6.0	3.0	3.8	2.0	3.8	21.0	10.6	56.9	25.7	17.6	17.8	NM	NM
ZSM uncatalogued [PT] *	THC245	F	Andramanolotra	38.7	12.6	15.1	5.9	3.2	3.7	2.5	3.7	22.3	11.0	59.6	25.9	18.2	18.9	NM	NM
ZSM uncatalogued [PT] *	THC355	F?	Andramanolotra	30.3	10.0	12.3	4.4	2.2	3.0	1.9	3.0	18.0	8.7	49.4	22.0	14.9	15.2	NM	NM
M. tricinctus
MNHN 1931.26 [LT]	NA	F	Befotaka	18.0	6.9	7.8	2.2	1.5	1.7	1.3	2.2	12.6	5.5	30.3	NM	9.1	NM	NA	NA
MNHN 1931.27 [PLT]	NA	M	Vondrozo	17.1	6.3	7.3	2.2	2.3	1.7	1.2	2.4	9.6	4.8	26.6	NM	8.2	NM	2.1	1.6
MNHN 1994.611 [PLT]	NA	M	Befotaka	17.1	6.4	7.1	2.2	2.2	1.6	1.4	2.2	10.0	5.0	26.1	NM	7.1	NM	2.3	1.5
MNHN 1994.612 [PLT]	NA	M	Befotaka	19.2	7.2	7.6	2.3	2.3	1.9	1.7	2.7	10.5	5.4	26.7	NM	8.7	NM	2.7	1.8
MNHN 1994.613 [PLT]	NA	M	Vondrozo	18.1	6.7	7.6	2.7	2.5	2.0	1.7	2.5	11.4	4.8	NM	NM	NM	NM	2.2	1.8
ZSM 177/2006	NA	F	Befotaka- Midongy	19.7	7.0	8.6	3.2	1.9	1.8	1.3	2.2	11.7	5.5	30.5	14.4	9.4	9.4	NA	NA
ZSM 2377/2007	ZCMV 5444	F	Manombo	23.4	8.1	10.2	3.3	2.4	2.9	1.9	2.7	14.4	6.6	38.8	17.8	10.9	11.9	NA	NA
ZSM 2415/2007	ZCMV 5420	M	Manombo	19.1	7.2	8.4	3.4	2.5	2.4	1.5	2.8	12.6	5.6	29.4	14.0	9.0	9.6	2.8	2.0
ZSM 176/2006	BOR 1066	M	Befotaka- Midongy	16.8	6.2	6.9	3.0	2.2	1.6	1.4	2.3	9.9	4.8	25.3	11.7	7.7	7.7	2.9	2.2
M. grubenmanni sp. nov.
ZSM 375/2006 [HT]	ZCMV 1404	M	An’Ala	17.9	6.5	7.6	3.0	2.8	2.4	1.2	2.0	11.4	4.8	28.3	13.3	8.8	8.7	3.0	1.9
ZFMK 62252 [PT]	NA	F	An’Ala	20.3	7.0	8.2	2.6	2.1	1.8	1.3	2.3	13.5	6.0	32.7	NM	10.0	NM	NA	NA
ZFMK 62253 [PT]	NA	F	An’Ala	19.2	6.5	7.7	2.3	2.0	1.8	1.3	2.5	12.7	5.5	30.8	NM	8.7	NM	NA	NA
ZFMK 62254 [PT]	NA	F	An’Ala	19.7	6.9	7.9	2.8	1.9	2.2	1.4	2.5	13.1	6.1	32.0	NM	10.4	NM	NA	NA
ZFMK 62251 [PT]	NA	M	An’Ala	16.7	6.0	6.7	2.5	2.2	1.8	1.4	2.0	11.1	4.8	27.4	NM	8.4	NM	2.5	1.8
ZFMK 62255 [PT]	NA	M	An’Ala	17.6	6.3	7.4	2.7	2.5	1.9	1.4	2.1	11.5	5.5	27.5	NM	9.0	NM	2.9	1.7
ZFMK 62256 [PT]	NA	M	An’Ala	17.8	6.5	7.3	2.6	2.4	1.6	1.3	2.1	11.7	5.5	29.4	NM	8.6	NM	2.6	1.7
ZFMK 62257 [PT]	NA	M	An’Ala	17.8	6.5	7.2	2.6	2.1	2.1	1.4	2.2	11.6	5.3	28.0	NM	8.6	NM	2.5	1.8
M. gudrunae sp. nov.
ZSM 146/2004 [HT]	FGZC 274	M	Manantantely	21.1	NM	8.8	3.2	2.8	2.5	1.7	2.6	13.0	5.6	32.9	14.4	9.7	10.0	4.4	2.5
ZSM 183/2021	ACZC 8516	F	Sainte Luce (S9)	28.6	11.1	11.7	4.4	2.9	2.8	2.0	3.3	17.8	8.2	44.9	20.2	13.6	13.4	NA	NA
ZSM 154/2004 [PT]	FGZC 286	F	Manantantely	24.4	10.1	8.9	3.8	2.5	2.5	1.9	3.1	8.5	6.6	NM	NM	10.8	12.5	1.4	1.1
ZSM 196/2005	FGZC 2594	F	Ste. Luce	26.3	10.4	9.5	4.2	2.6	2.7	1.7	2.6	10.6	7.5	NM	NM	12.6	13.7	NA	NA
ZSM 68/2004 [PT]	FGZC 115	F	Andohahela	25.3	10.5	10.4	3.8	2.4	2.5	2.0	2.9	9.8	8.7	NM	NM	12.9	14.3	1.5	1.1
ZSM 96/2004 [PT]	FGZC 168	F	Andohahela	26.5	10.8	10.1	3.9	3.1	2.9	2.2	2.9	10.7	8.2	NM	NM	12.9	14.0	1.5	1.0
MRSN A7046	FAZC 15427	F	Tsitongambarika	22.6	8.0	9.6	3.2	2.2	2.1	2.0	2.8	13.4	6.7	36.3	16.5	11.0	10.7	NA	NA
ZSM 182/2021	ACZC 8515	M	Sainte Luce (S9)	24.4	9.0	10.1	2.7	2.6	2.7	1.6	2.9	14.4	6.8	36.3	16.3	11.1	11.3	3.2	2.5
ZSM 136/2004 [PT]	FGZC 250	M	Manantantely	21.8	8.6	9.4	4.0	3.3	2.4	1.4	2.7	14.6	6.8	34.9	16.4	10.9	10.8	3.7	2.8
ZSM 138/2004 [PT]	FGZC 259	M	Manantantely	20.5	7.9	9.0	3.8	2.9	2.0	1.4	2.6	13.7	5.8	31.8	14.2	9.8	9.3	4.2	2.5
ZSM 95/2004 [PT]	FGZC 167	M	Andohahela	20.2	8.6	7.8	2.9	2.7	2.1	1.7	2.3	8.5	6.2	NM	NM	9.7	10.5	2.3	1.6
MRSN A7044	FAZC 15282	M	Sainte Luce	23.0	8.0	9.6	3.6	2.7	2.6	1.4	2.9	13.6	6.5	33.7	15.2	10.4	11.0	3.2	2.1
MRSN A7045	FAZC 15419	M	Tsitongambarika	21.9	8.0	9.2	3.7	3.0	2.4	1.8	2.8	12.6	6.4	34.5	15.4	10.0	10.2	2.5	2.3
Mantidactylus biporus
BMNH 1947.2.26.47 [LT*]	NA	F	Madagascar	35.8	13.7	13.9	4.7	2.8	2.8	1.7	3.7	18.1	8.5	51.2	22.3	15.3	NM	NA	NA
BMNH 1947.2.26.48 # [PLT]	NA	F?	Madagascar	28.0	12.1	10.0	4.6	2.3	2.8	1.9	3.5	9.4	8.6	NM	NM	12.2	12.8	1.5	1.2
BMNH 1947.2.26.50 # [PLT]	NA	F?	Madagascar	29.4	11.3	9.7	4.2	2.5	2.4	1.8	3.3	10.0	7.2	NM	NM	12.6	12.0	1.4	1.2
BMNH 1947.2.26.49 [PLT]	NA	M	Madagascar	29.9	10.7	11.9	4.1	3.1	2.7	2.2	2.7	15.9	7.8	41.7	19.5	13.6	NM	3.9	2.5
MRSN A6180	FAZC 13480	M	Betampona	31.6	12.6	12.5	4.8	3.3	2.7	2.6	3.6	17.4	8.7	43.3	20.1	14.0	12.8	3.0	2.2
MRSN A6266	FAZC 13675	M	Betampona	29.9	12.0	12.1	4.9	3.7	2.9	2.1	3.7	16.6	8.0	41.5	19.4	14.0	11.5	3.3	2.1
MRSN A6374	FAZC 13835	M	Betampona	31.0	12.6	12.7	4.5	3.4	2.7	2.2	3.3	17.6	8.4	42.0	19.6	14.1	12.5	3.4	2.7
ZSM 1982/2006	ZCMV 2425	M	An’Ala	28.0	11.0	11.0	3.8	3.0	2.7	1.7	2.7	15.1	7.0	38.2	17.3	11.8	11.2	2.3	1.8
ZSM 396/2006	ZCMV 1483	F	An’Ala	30.5	11.5	11.4	4.3	3.0	2.8	1.7	2.9	16.0	8.0	42.9	18.7	13.2	12.1	1.4	1.2
ZSM 184/2021	ACZCV 201	F?	Betampona: Vohitsivalana	19.3	8.0	8.6	2.8	1.8	2.4	1.8	2.7	11.0	5.7	29.2	13.2	8.7	8.6	NA	NA
Mantidactylus augustini sp. nov. (Ca22)
ZSM 122/2002 [HT]	FGMV 2001.1388 (2002. A4)	M	Andranofotsy	23.9	10.1	10.3	3.9	3.2	2.5	1.6	2.8	15.9	6.9	36.7	16.9	11.7	10.5	3.3	2.0
MRSN A3600 [PT]	FN 7678 (ACZC 4904)	F	Masoala forest	25.0	9.6	10.3	3.7	3.4	1.6	1.6	3.0	14.8	7.5	38.3	17.0	11.8	10.6	NA	NA
ZFMK 70481 #	NA	F	Masoala	24.2	9.5	9.0	3.8	2.6	2.0	1.8	3.0	9.7	6.8	NM	NM	10.4	11.4	1.4	1.3
MRSN A2905 [PT]	FN 7238 (ACZC 4897)	F?	Masoala forest	24.4	10.3	10.8	4.0	2.9	2.7	1.7	3.0	NM	7.4	NM	17.4	12.6	11.1	NA	NA
MRSN A3540 [PT]	ACZC 4898	F?	Masoala forest	22.6	9.3	10.0	3.7	2.4	2.4	1.6	2.9	14.8	6.9	37.8	17.4	11.8	11.1	NA	NA
ZSM 740/2009 [PT]	ZCMV 11170	F	Melivinany, Manompana	20.9	8.2	8.6	3.1	2.7	1.9	1.7	2.9	13.9	6.6	33.0	15.3	10.9	9.8	1.7	1.3
MRSN A6740 [PT]	FAZC 14292 (ACZC 4906)	FJ?	Farankaraina	16.4	NM	NM	NM	NM	NM	NM	NM	NM	NM	NM	NM	NM	NM	NA	NA
MRSN A3737 [PT]	FAZC 10009 (ACZC 4905)	J	Masoala forest	16.4	NM	NM	NM	NM	NM	NM	NM	NM	NM	NM	NM	NM	NM	NA	NA
Mantidactylus bletzae sp. nov.
ZSM 829/2014 [HT]	ZCMV 14771	F	Ivohibe	27.3	10.6	10.5	3.4	2.4	1.8	1.8	3.0	16.1	7.9	41.0	18.6	12.2	12.4	NA	NA
ZSM 827/2014 [PT]	ZCMV 14763	F	Ivohibe	25.8	9.8	9.9	3.4	2.6	2.0	1.6	2.9	15.5	7.4	39.3	18.1	12.2	11.8	NA	NA
ZSM 828/2014 [PT]	ZCMV 14768	F	Ivohibe	26.2	9.8	10.1	3.5	2.3	2.1	1.7	2.6	15.6	8.3	40.5	18.6	12.3	12.0	NA	NA
Mantidactylus brevirostris sp. nov.
MRSN A6257 [HT]	FAZC 13581	M	Betampona: Sahamben- drana	23.2	8.8	9.3	3.5	2.7	2.4	1.6	3.4	14.7	6.5	34.9	15.6	10.7	10.4	2.7	1.6
ZSM 185/2021 [PT]	ACZCV 265	F	Betampona: Sahabefoza	28.4	11.3	11.4	3.8	2.6	2.4	1.6	2.8	16.3	7.3	40.6	17.9	12.0	12.0	NA	NA
Mantidactylus eulenbergeri sp. nov. (Ca23)
ZSM 85/2002 [HT]	FGMV 2001.1092	M	Andasibe	21.6	9.2	8.7	3.3	3.0	1.8	1.4	2.3	12.5	5.7	29.7	13.2	9.0	7.9	3.4	2.3
ZFMK 62214 #	NA	M	Andasibe	23.3	9.1	8.1	3.7	2.7	2.1	1.8	2.5	9.2	6.6	NM	NM	10.1	10.1	2.2	2.2
MRSN A7047 [PT]	FAZC 15517	M	Maromizaha	21.9	9.5	9.5	3.5	3.1	2.1	1.7	2.9	13.9	6.4	32.4	14.8	10.0	9.6	2.5	1.5
MRSN A7048 [PT]	FAZC 15549	M	Maromizaha	20.9	8.1	8.6	3.0	2.5	1.8	1.8	2.7	12.0	6.3	31.2	13.9	9.7	9.4	2.0	1.4
ZSM 199/2021 [PT]	FAZC 15516	M	Maromizaha	20.5	8.8	9.2	3.3	2.5	1.8	1.7	3.0	12.6	6.5	30.3	13.7	9.5	9.4	2.7	1.6
MRSN A6700 [PT]	RJS 1820 (ACZC 4200)	F	Anivorano Est forest	26.4	11.1	12.3	4.4	2.9	2.5	2.0	3.4	16.2	7.6	39.3	17.8	12.0	12.0	NA	NA
ZSM 84/2002 [PT]	FGMV 2001.1090	F	Andasibe	28.0	10.5	10.7	3.5	2.6	2.1	1.9	2.9	15.6	7.4	39.0	17.1	11.9	11.4	2.3	1.0
ZSM 198/2021 [PT]	FAZC 15509	F	Maromizaha	26.7	10.4	11.0	3.4	2.4	2.1	2.0	2.5	16.0	7.9	35.6	16.0	10.0	10.4	NA	NA
MRSN A7049 [PT]	FAZC 15540	F	Maromizaha	25.0	10.1	11.0	3.6	2.5	1.9	1.9	3.0	14.1	7.2	37.4	16.8	11.6	11.0	NA	NA
ZSM 919/2003 [PT]	FGMV 2002.949	F	Vohidrazana	20.0	8.5	8.5	3.2	2.3	1.9	1.1	2.3	12.6	5.8	29.4	13.9	9.3	9.0	1.4	1.2
Mantidactylus glosi sp. nov.
ZSM 434/2006 [HT]	ZCMV 3366	M	Ranomafana	21.0	8.7	8.6	3.2	2.8	2.1	1.4	2.3	11.9	5.6	28.9	13.5	9.3	9.1	2.5	1.5
ZSM 433/2006 [PT]	ZCMV 3364	F	Ranomafana	25.1	9.7	10.0	3.4	3.1	2.3	1.4	2.5	13.8	6.5	34.1	16.0	11.3	10.1	1.2	1.0
Mantidactylus stel- liger sp. nov.
ZSM 2381/2007 [HT]	ZCMV 5932	M	Ambohitsara	22.6	9.7	10.0	3.9	2.6	2.1	1.5	2.6	13.8	6.4	33.1	15.0	9.8	9.9	2.8	2.2
ZSM 2380/2007 [PT]	ZCMV 5922	M	Ambohitsara	21.6	8.8	9.4	3.4	2.5	2.0	1.6	2.8	13.8	6.4	33.3	14.9	10.2	10.1	2.6	1.9
ZSM 2379/2007 [PT]	ZCMV 5870	F	Ambohitsara	31.1	13.0	13.2	4.6	3.4	2.8	2.6	3.0	19.1	9.3	46.2	20.8	14.5	13.6	1.1	0.8
Mantidactylus inaudax bona species (Ca17)
MZUT An727.1 [LT*]	NA	F		32.1	12.7	12.4	4.9	3.5	2.2	2.0	3.3	17.7	8.5	47.9	21.8	15.4	13.5	NA	NA
ZSM 180/2005	FGZC 2146	M	Ambohitantely	26.2	10.2	10.5	3.7	4.1	1.9	1.8	2.6	14.1	6.8	34.4	16.6	11.8	10.2	4.5	1.7
ZFMK 60142 #	NA	F	Ambohitantely	26.9	11.8	9.4	3.8	2.6	2.2	2.1	3.2	9.1	7.8	NM	NM	11.0	11.0	NA	NA
ZMA 19310 #	FGMV 2002.2435	F	Fierenana	33.1	13.3	12.7	5.2	3.3	2.7	2.1	4.1	11.3	9.1	NM	NM	14.6	14.6	1.6	1.3
ZMA 19311 #	FGMV 2002.2439	F	Fierenana (pitfall forest)	27.4	11.3	9.9	3.8	3.1	2.3	1.9	3.3	11.4	8.6	NM	NM	13.8	13.6	1.6	1.4
ZMA 19314 #	FGMV 2002.2429	F	Fierenana (Camp)	27.5	11.5	9.9	4.0	3.0	2.1	1.7	3.0	10.9	8.6	NM	NM	14.0	14.5	1.2	1.2
ZMA 19331 #	FGMV 2002.2441	F	Fierenana (pitfall forest)	27.0	11.1	9.6	3.8	3.0	2.1	1.9	3.2	11.4	8.8	NM	NM	13.6	13.7	1.5	1.2
ZSM 181/2005	FGZC 2147	F	Ambohitantely	29.4	11.0	11.2	4.0	3.1	2.3	1.8	2.8	15.5	7.5	40.2	19.3	13.1	11.6	NA	NA
ZMA 19341 #	FGMV 2002.2252	F?	Fierenana (Camp)	33.6	12.9	11.5	4.7	3.6	2.4	1.9	3.8	11.9	9.9	NM	NM	15.7	15.4	1.7	1.2
ZFMK 60141 #	NA	M	Ambohitantely	21.7	9.8	7.8	3.2	2.9	1.7	1.7	2.5	8.5	6.4	NM	NM	9.7	10.1	2.2	1.6
ZMA 19343 #	FGMV 2002.2423	M	Fierenana (Camp)	27.4	11.0	9.6	3.7	3.4	2.1	1.8	3.3	10.0	8.7	NM	NM	12.5	12.3	2.7	2.0
ZMA 19345 #	FGMV 2002.2251	M	Fierenana (Camp)	25.8	10.5	9.4	3.8	3.2	2.1	1.7	3.0	9.5	7.6	NM	NM	11.3	11.0	3.0	2.2
ZSM 1768/2008	ZCMV 8869	M	Forest fragment Brieville	25.7	10.6	10.5	4.0	3.6	2.3	1.8	3.2	15.0	6.7	38.2	17.6	12.1	10.4	2.7	1.9
ZSM 363/2010	FGZC 4358	M	Fanamby Forest, Anjozorobe	23.7	9.0	9.1	3.4	3.4	2.2	1.6	2.3	13.0	6.8	31.6	15.2	10.7	9.4	2.9	2.1
ZSM 398/2006	ZCMV 3259	M	Moramanga- Anosibeanala	30.0	10.7	11.9	4.3	3.6	2.3	2.1	3.3	15.8	8.3	42.2	20.7	14.5	11.5	2.8	2.2
ZSM 548/2009	ZCMV 11213	M	Makira West	22.7	8.7	9.5	3.8	3.3	2.2	1.5	2.4	13.5	7.0	33.9	16.5	11.7	9.8	4.4	2.0
ZSM 85/2016	MSZC 0161	M	Ampotsidy	29.1	11.2	12.2	4.2	3.8	2.3	2.2	3.4	15.9	8.2	40.1	19.2	13.5	11.4	3.2	3.0
ZSM 88/2016	MSZC 0178	M	Ampotsidy	28.4	10.4	11.4	4.2	3.9	2.1	1.7	3.0	15.5	8.0	38.3	18.2	12.9	11.0	5.3	2.2
Mantidactylus manerana manerana ssp. nov. (Ca16)
ZSM 500/2016 [HT]	ZCMV 15162	M	Marojejy	27.5	10.2	11.0	3.9	3.3	2.8	1.8	3.0	16.0	8.5	40.7	18.6	12.9	12.0
ZSM 206/2005 [PT]	FGZC 2837	F	Marojejy Camp Simpona	29.4	10.7	11.5	4.3	2.6	2.5	1.9	2.8	17.4	7.7	44.0	19.3	13.8	12.9	NA	NA
ZSM 5084/2005 [PT]	ZCMV 2019	F	Marojejy Camp Simpona	28.0	11.2	11.0	3.6	2.8	2.7	1.8	2.8	17.6	8.1	45.2	20.3	14.8	13.4	1.2	1.2
ZSM 207/2005 [PT]	FGZC 2846	M	Marojejy Camp Simpona	26.0	10.3	10.3	4.0	3.2	2.5	1.7	2.6	14.9	6.3	37.5	17.1	11.9	10.8	3.4	2.2
ZSM 208/2005 [PT]	FGZC 2847	M	Marojejy Camp Simpona	26.8	10.4	10.8	3.5	3.0	2.4	1.9	3.0	16.3	7.7	41.2	18.3	13.0	11.6	3.3	2.0
ZSM 209/2005 [PT]	FGZC 2881	M	Marojejy Camp Simpona	25.0	9.4	9.4	3.3	2.2	2.1	2.0	2.7	15.8	7.2	37.6	16.8	11.7	11.4	3.6	1.9
Mantidactylus manerana fotaka ssp. nov.
ZSM 1588/2012 [HT]	FGZC 3776	M	Sorata	26.2	10.2	10.3	3.8	3.5	2.5	1.8	2.8	13.7	6.9	36.8	17.5	12.1	10.5	4.6	2.3
ZSM 1587/2012 [PT]	FGZC 3770	F	Sorata	29.2	11.4	11.3	3.9	2.7	2.3	1.8	2.8	15.7	7.8	39.5	18.7	12.7	11.6	1.4	1.0
Mantidactylus manerana antsanga ssp. nov.
MRSN A5633 [HT]	FAZC 13371 (ACZC 10079)	F	Sahavontsira	37.5	13.0	14.1	5.2	3.6	2.7	2.6	3.7	21.3	9.6	53.0	25.1	17.3	15.8	NA	NA