An inordinate fondness for inconspicuous brown frogs: integration of phylogenomics, archival DNA analysis, morphology, and bioacoustics yields 24 new taxa in the subgenus Brygoomantis (genus Mantidactylus) from Madagascar Author Scherz, Mark D. Author Crottini, Angelica CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal Author Hutter, Carl R. 0000-0001-6381-6339 Museum of Natural Sciences and Department of Biological Sciences, Louisiana State University, Baton Rouge, LA 70803, USA Ễcarl. hutter @ gmail. com; https: // orcid. org / 0000 - 0001 - 6381 - 6339 USAỄcarl.hutter@gmail.com Author Hildenbrand, Andrea Hauptstr. 13, 82234 Wessling, Germany Ễandrea @ gutachten-hildenbrand. de GermanyỄandrea@gutachten-hildenbrand.de Author Andreone, Franco Museo Regionale di Scienze Naturali, Via G. Giolitti, 36, 10123 Torino, Italy Author Fulgence, Thio Rosin 0000-0001-7205-7282 Natural and Environmental Sciences, Regional University Centre of the SAVA Region (CURSA), Antalaha, Madagascar Ễthiorosinf @ yahoo. fr; https: // orcid. org / 0000 - 0001 - 7205 - 7282 MadagascarỄthiorosinf@yahoo.fr Author Köhler, Gunther Author Ndriantsoa, Serge Herilala Author Ohler, Annemarie Author Preick, Michaela Author Rakotoarison, Andolalao Mention Zoologie et Biodiversité Animale, Université d’Antananarivo, BP 906, Antananarivo, 101 Madagascar Author Rancilhac, Loïs Author Raselimanana, Achille P. Mention Zoologie et Biodiversité Animale, Université d’Antananarivo, BP 906, Antananarivo, 101 Madagascar Author Riemann, Jana C. Author Rödel, Mark-Oliver Author Rosa, Gonçalo M. Author Streicher, Jeffrey W. Author Vieites, David R. Author Köhler, Jörn Author Hofreiter, Michael Author Glaw, Frank Author Vences, Miguel text Megataxa 2022 2022-12-15 7 2 113 311 http://dx.doi.org/10.5281/zenodo.6687412 journal article 217510 10.11646/megataxa.7.2.1 8af27bde-abaa-4f0a-8f84-8b6cc8a97034 2703-3090 7441023 2FD8C310-6486-4592-92F6-5EB894EBD6AC
TABLE 3. List of localities mentioned in the text, geographical coordinates, elevations, and species of Mantidactylus (Brygoomantis) occurring at the respective sites. Elevation (m above sea level [a.s.l.]) is partly based on references, own GPS readings, or has been inferred via GIS from the coordinates of records (marked with an asterisk). When elevation is given as NA (not applicable), it is because specimens were collected in that general area, but without precise coordinates from which to extract elevational range of its occurrence. Note that many additional (identified and unidentified) records of Brygoomantis species are available from published herpetological surveys (Rakotondravony & Goodman 2011) including numerous protected areas (e.g. Raselimanana et al. 2018), but since reliable species identification is almost impossible without examining specimens and/or genetic data, they are not considered in this table.
LocationLat.Lon.Elev.CladeSpecies
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20 km north of Vatomandry-19.192648.912815*fergusoniM. georgei
Alaotra region (swamp)-17.77448.083933*betsileanusM. betsileanus
curtusM. alutus
Ambahavala-24.142747.1057346tricinctusM. tricinctus
Ambatobe-15.2550.4314fergusoniM. fergusoni
inaudaxM. manerana antsanga
tricinctusM. grubenmanni
Ambatomandondona-18.916348.4196932*betsileanusM. katae
Ambatovaky-21.282647.32731201betsileanusM. betsileanus
Ambinanifaho-14.616750.133327*fergusoniM. fergusoni
Ambodiriana-16.674649.7028158fergusoniM. fergusoni
Ambodisakoa-17.31248.6661804betsileanusM. betsileanus
Ambodivohitra-14.840949.9524411*betsileanusM. jonasi
Ambohimitombo-20.7247.431150curtusM. ambohimitombi ambohimitombi
Ambohitantely-18.17247.3021520betsileanusM. betsileanus
curtusM. alutus
curtusM. ambohimitombi miloko
inaudaxM. inaudax
Ambohitantely: Jardin botanique-18.171747.28171580betsileanusM. betsileanus
curtusM. ambohimitombi miloko
inaudaxM. inaudax
Ambohitantely: Site 2-18.178347.29041546betsileanusM. betsileanus
inaudaxM. inaudax
Ambohitantely: Zone 3-18.199547.28091574*betsileanusM. betsileanus
Ambohitsara-21.357247.8157294betsileanusM. katae
stelligerM. stelliger
tricinctusM. grubenmanni
Ambositra-20.516747.251300curtusM. curtus
Ampasimazava-16.922749.2493445fergusoniM. fergusoni
Ampofoko (old camp)-15.422949.12091034betsileanusM. jonasi
Ampotsidy-14.413548.71731227– 1380betsileanusM. jonasi
inaudaxM. inaudax
An’Ala-18.9348.47840betsileanusM. katae
biporusM. biporus
tricinctusM. grubenmanni
Andapa-14.633349.6167580ulcerosusM. bellyi
Andasibe: orchid garden-18.932748.4132920betsileanusM. betsileanus
Andasibe: stream along road-18.92948.413920betsileanusM. katae
Andasibe/Analamazaotra-18.933348.4167920betsileanusM. betsileanus
betsileanusM. katae
biporusM. eulenbergeri
Andohahela: Camp 2005-24.544046.71411548betsileanusM. kortei
Andohahela: Camp 1-24.758646.8542247betsileanusM. katae
betsileanusM. tripunctatus
tricinctusM. gudrunae
Andrakata-14.616749.7167460ulcerosusM. bellyi
Andramanolotra-13.996950.096316*fergusoniM. marintsoai
Andrangoloaka-19.0047.951380curtusM. alutus
Andranofotsy-15.435349.843985biporusM. augustini
fergusoniM. fergusoni
Andranonafindra Forest-14.73648.54831180betsileanusM. jonasi
Andringitra-22.246.9NAbetsileanusM. katae
curtusM. bourgati
curtusM. madecassus
Andringitra: Andohariana Plateau-22.180346.90032030curtusM. bourgati
Andringitra eastern slopes: Imaitso-22.140346.94691509curtusM. bourgati
Andringitra western slopes: Siranandambo-22.130346.84781590curtusM. bourgati
Andringitra western slopes: Iantaranomby-22.129046.84671580curtusM. bourgati
Andringitra western slopes: Andramena-22.127346.85331740curtusM. bourgati
Andringitra eastern slopes: Asaramanitra-22.135846.88901590curtusM. bourgati
Andringitra eastern slopes: Riandahy-22.145846.89151730curtusM. bourgati
Andringitra: Ambalamarovandana-22.22646.9341530curtusM. bourgati
Andringitra: Cuvette Boby-22.194746.88972488curtusM. bourgati
curtusM. madecassus
Andringitra: Pic Boby-22.246.92NAcurtusM. madecassus
Angavokely-18.916747.73331640curtusM. ambohimitombi miloko
Angorony: fragment near Maromandia-14.221148.1421115ulcerosusM. ulcerosus
Anivorano Est: Andrarihitra,-18.763848.946860betsileanusM. incognitus
VohiposabiporusM. eulenbergeri
fergusoniM. georgei
Anjozorobe-18.447.86671250betsileanusM. betsileanus
inaudaxM. inaudax
Ankarafantsika-16.283946.7749853*ulcerosusM. ulcerosus
Ankarafantsika: Ampondrabe Forest-16.32546.9233250ulcerosusM. ulcerosus
Ankarafantsika: Andasiravina Forest-16.303346.93150ulcerosusM. ulcerosus
Ankarana-12.933649.126990ulcerosusM. bellyi
Ankaratra-19.333347.2667NAcurtusM. alutus
curtusM. ambohimitombi ambohimitombi
curtusM. pauliani
Ankaratra: Analafohy-19.344247.2752146curtusM. ambohimitombi ambohimitombi
Ankaratra: forest Ambohimiradrana road-19.332547.27092000curtusM. ambohimitombi ambohimitombi
Ankaratra: Tavolotara-19.345847.27912019curtusM. ambohimitombi ambohimitombi
Ankaratra: Tsiafajavona plateau-19.328347.26172380curtusM. ambohimitombi ambohimitombi
Ankazomivady-20.772247.18771735curtusM. ambohimitombi miloko
curtusM. curtus
Anosibe An’Ala-19.433348.2167636betsileanusM. betsileanus
Antambato: swamp ca 4 km from Antambato village (Antsaloana)-14.481148.89991258betsileanusM. jonasi
Antambato: village stream-14.49348.86861188betsileanusM. jonasi
Antanambe-16.429949.7846263fergusoniM. fergusoni
Antara-16.887549.1832481*betsileanusM. betsileanus
Antoetra-20.801347.36471680curtusM. alutus
curtusM. ambohimitombi ambohimitombi
curtusM. curtus
Antoetra: Farimazava Forest-20.83547.33251380– 1420curtusM. ambohimitombi ambohimitombi
curtusM. curtus
Antokotelo-18.427749.0096308*fergusoniM. georgei
Antsahanoro-14.84550.133666*fergusoniM. fergusoni
Antsatramidola-15.63448.9675404ulcerosusM. ulcerosus
Antsirakambiaty forest-20.594946.56381601betsileanusM. betsileanus
curtusM. curtus
Bealanana-Antsohihy forest fragment-14.721548.56271187betsileanusM. jonasi
Beanka-18.023644.5022254curtusM. mahery
Befanjana-16.6849.59NAbetsileanusM. betsileanus
biporusM. augustini
fergusoniM. fergusoni
inaudaxM. manerana antsanga
tricinctusM. grubenmanni
Befotaka-Midongy: Rozabe Forest-23.736847.0227780–900tricinctusM. tricinctus
Belambo-14.548749.7489234*fergusoniM. fergusoni
ulcerosusM. bellyi
Bemandrevo/Samangorona/ Alaotra/Mangoro-18.40548.63661*betsileanusM. incognitus
Bemanevika-14.25948.7781130betsileanusM. jonasi
Bemanevika: Camp 1, Antsirakala-14.430648.60181468betsileanusM. jonasi
Bemanevika: Camp 2-14.359948.59021538betsileanusM. jonasi
Bemanevika: River-14.482548.62721109betsileanusM. jonasi
Bemanevika/Antsirabe-Nord-13.986449.951961*fergusoniM. marintsoai
Benavony-13.748.4833140ulcerosusM. ulcerosus
Berara-14.309247.9153170ulcerosusM. ulcerosus
Betampona-17.888349.2277190–517betsileanusM. betsileanus
biporusM. biporus
biporusM. brevirostris
fergusoniM. georgei
tricinctusM. grubenmanni
Betampona: Maintimbato-17.894049.2283NAbetsileanusM. georgei
Betampona: Rendrirendry-17.918649.2103325betsileanusM. betsileanus M. georgei
Betampona: Sahabefoza-17.915249.2090349betsileanusM. betsileanus
biporusM. brevirostris
Betampona: Sahaindrana-17.8938349.19972327betsileanusM. betsileanus
biporusM. biporus
Betampona: Sahambendrana-17.898449.2154458biporusM. brevirostris
Betampona: Vohitsivalana-17.884649.2008517biporusM. biporus
fergusoniM. georgei
Bevitagnono Forest-14.738648.51721024– 1041ulcerosusM. ulcerosus
Camp Ambatofotsy-19.543148.3165907betsileanusM. katae
Col des Tapias-20.2347.081503curtusM. curtus
Commune d’Andekaleka: Marovato-18.68648.6055787betsileanusM. incognitus
Fanambana-13.613850.001953ulcerosusM. bellyi
Fierenana (Camp)-18.543348.4489948betsileanusM. betsileanus
inaudaxM. inaudax
Fivahona: Ambavala-22.044746.90141508betsileanusM. betsileanus
curtusM. bourgati
Fivahona: Velotsoa-22.070446.87591278betsileanusM. betsileanus
curtusM. bourgati
forest fragment, road from Lake Alaotra to Brieville (Andranogorika)-17.767847.98421147betsileanus inaudaxM. betsileanus M. inaudax
Ibity-20.116747.01672090curtusM. alutus
curtusM. curtus
Ifanadiana-21.347.6333530betsileanusM. katae
Irogno Forest-14.7548.492958ulcerosusM. ulcerosus
Isalo-22.4245.27NAbetsileanusM. noralottae
betsileanusM. riparius
curtusM. mahery
Isalo: Ambovo-22.299845.3526996betsileanusM. noralottae
Isalo: Andriamanero-22.3545.4640betsileanusM. noralottae
betsileanusM. riparius
Isalo: Anjofo-22.369445.361800betsileanusM. noralottae
Isalo: Canyon des rats-22.480345.378960curtusM. mahery
Isalo: Cascade des Nymphes-22.537345.3759870betsileanusM. riparius
curtusM. mahery
Isalo: Forêt d’Analalava-22.587945.1308719curtusM. mahery
Isalo: Hotel Isalo Ranch-22.592645.3928804curtusM. mahery
Isalo: Namazaha-22.536745.3748871betsileanusM. riparius
curtusM. mahery
Isalo: Oasis-22.626945.3533776curtusM. mahery
Isalo: Piscine naturelle-22.566445.364920betsileanusM. riparius
Isalo: Zahavola-22.803745.2458850curtusM. mahery
Itremo-20.509146.49841648betsileanusM. betsileanus
curtusM. ambohimitombi marefo
curtusM. curtus
Ivohibe: Camp 3 (high elevation)-22.497146.95761575biporusM. bletzae
Ivoloina-18.066749.425fergusoniM. georgei
km 27 on RN Antsohihy- Mandritsara-15.053248.2064140ulcerosusM. ulcerosus
Lac Alaotra-17.5948.52850betsileanusM. betsileanus
Lokobe National Park-13.398848.31830–110ulcerosusM. ulcerosus
Mahanoro-19.906248.815610betsileanusM. incognitus
Mahasoa campsite-17.297748.7021032betsileanusM. betsileanus
Mahatsara-Mantadia/Mantady-18.828848.4327995betsileanusM. katae
Makay-21.686645.1700174curtusM. mahery
Makira western slope: Camp 0,-15.488949.0785603betsileanusM. jonasi
SahaovycurtusM. mahery
ulcerosusM. ulcerosus
Makira western slope: Camp 1, Angozongahy-15.43749.11861009betsileanusM. jonasi
Makira western slope: Hevirina-15.44949.11191093inaudaxM. inaudax
(Pandanus swamp)ulcerosusM. ulcerosus
Makira western slope: Satellite Camp, source of Fotsialanana river-15.466849.12891067betsileanusM. jonasi
Makira: Ambodivohangy-15.289949.620383fergusoniM. fergusoni
Manantantely-24.98346.917180betsileanusM. tripunctatus
tricinctusM. gudrunae
Mandena-24.952247.00398betsileanusM. tripunctatus
Mandraka-18.928947.89361210betsileanusM. betsileanus
Mangindrano: Camp 0 near Mangindrano (Ambinanitelo)-14.225448.96341171betsileanusM. jonasi
Manombo-23.0147.733430tricinctusM. tricinctus
Manombo: Camp-23.028347.731544tricinctusM. tricinctus
Manongarivo: Camp 0-13.975648.4267688ulcerosusM. schulzi
Manongarivo: Camp 1-13.976948.4219751ulcerosusM. steinfartzi
Mantasoa-19.016747.83331430curtusM. alutus
Mariavaratra-21.624547.41941144betsileanusM. katae
Maroantsetra-15.42749.7415170fergusoniM. georgei
Marojejy-14.406749.7671NAfergusoniM. fergusoni
Marojejy: above Camp Simpona-14.440849.73991576inaudaxM. manerana manerana
Marojejy: between camps 2 and 3-14.434449.7666615*fergusoniM. fergusoni
inaudaxM. manerana manerana
Marojejy: Camp 0-14.446349.7852310fergusoniM. fergusoni
Marojejy: Camp 1 (Mantella)-14.437749.7756481fergusoniM. fergusoni
Marojejy: Camp 3 (Simpona)-14.436749.74341326betsileanusM. jonasi
fergusoniM. fergusoni
inaudaxM. manerana manerana
Marojejy: close to Camp 1 (Mantella)-14.439449.7771465*fergusoniM. fergusoni
Marojejy: east of tourist trail-14.44649.8234273*fergusoniM. fergusoni
fergusoniM. marintsoai
ulcerosusM. bellyi
Marolambo-20.0548.1167490betsileanusM. katae
Marolambo: Ambodisavoka-20.091948.3222287betsileanusM. katae
Maromizaha-18.976248.4648980–1100betsileanusM. betsileanus
betsileanusM. katae
biporusM. eulenbergeri
Marotandrano-Riamalandy-16.283348.8144815–1015betsileanusM. betsileanus
Marovitsika-22.52546.98331000betsileanusM. katae
Masoala-15.5550.1167NAbetsileanusM. jonasi
biporusM. augustini
fergusoniM. fergusoni
Masoala: Ilampy Corridor15.392050.0470ca 550biporusM. augustini
Masoala: Ambatoledama Corridor-15.26749.983ca 1000biporusM. augustini
Midongy du Sud-23.583347.0167600tricinctusM. tricinctus
Montagne d’Ambre: Gîte d’Etape-12.526849.17211050betsileanusM. jonasi
ulcerosusM. bellyi
Montagne d’Ambre: Andrano créole-12.49549.1848751–772ulcerosusM. bellyi
Montagne d’Ambre: Antomboko-12.490749.1716652–730ulcerosusM. bellyi
Montagne d’Ambre: Antsakomboiny-12.468849.2207467–480ulcerosusM. bellyi
Montagne d’Ambre: Betsikoboko-12.490849.1787623–743ulcerosusM. bellyi
Montagne d’Ambre: Cascade Antakarana-12.520649.1724997–1061ulcerosusM. bellyi
Montagne d’Ambre: Cascade sacrée-12.528649.17081057– 1192ulcerosusM. bellyi
Montagne d’Ambre: high-12.596349.15921372betsileanusM. jonasi
elevation around Grand LaculcerosusM. bellyi
Montagne d’Ambre: high elevation near Lac Maudit-12.583949.15331225– 1274betsileanusM. jonasi
ulcerosusM. bellyi
Montagne d’Ambre: low elevation-12.516549.1713484–737ulcerosusM. bellyi
Montagne d’Ambre: mid-elevation-12.556749.16231028– 1105betsileanusM. jonasi
ulcerosusM. bellyi
Montagne d’Ambre: RE1-12.533449.18771010– 1057ulcerosusM. bellyi
Montagne d’Ambre: RW1-12.52749.16761089– 1125betsileanusM. jonasi
ulcerosusM. bellyi
Montagne d’Ambre: RW2-12.520949.16661042– 1093ulcerosusM. bellyi
Montagne d’Ambre: West Slope-12.588349.128945–966ulcerosusM. bellyi
Montagne d’Ambre: Zanakatomboko-12.48749.1701670–715ulcerosusM. bellyi
Montagne des Français: Andavakoera-12.333349.3580–200ulcerosusM. bellyi
Moramanga-Anosibe An’Ala/ Besariaka crossroad-19.099448.2453972*inaudaxM. inaudax
Nahampoana-24.979446.983916betsileanusM. tripunctatus
Namoly: Analabe-22.105246.94461626*betsileanusM. betsileanus
curtusM. bourgati
Nosy Be-13.350648.275919ulcerosusM. ulcerosus
Nosy Boraha-16.835249.9271NAfergusoniM. jahnarum
Nosy Boraha: Maromandia village-16.908949.867820fergusoniM. jahnarum
Nosy Mangabe-15.489349.76450–115fergusoniM. fergusoni
Pic Saint-Louis-25.00846.962365betsileanusM. tripunctatus
Ranomafana-21.2547.45930betsileanusM. betsileanus
betsileanusM. katae
biporusM. glosi
Ranomafana: Ambatolahidimy-21.247147.4190984betsileanusM. katae
Ranomafana: Ambatolahy-21.243947.4262950betsileanusM. betsileanus
betsileanusM. katae
Ranomafana: Ambatovory-21.238047.4248966betsileanusM. katae
biporusM. glosi
Ranomafana: Ambolo I-21.263147.5070650betsileanusM. betsileanus
betsileanusM. katae
tricinctusM. grubenmanni
Ranomafana: Ambolo II-21.263947.5086675betsileanusM. betsileanus
betsileanusM. katae
Ranomafana: Andalangina-21.298647.6022486betsileanusM. betsileanus
betsileanusM. katae
biporusM. glosi
Ranomafana: Beremby,-21.231047.5065640betsileanusM. katae
AntaramanavanatricinctusM. grubenmanni
Ranomafana: Beremby, Beremby-21.240447.5250618betsileanusM. betsileanus
betsileanusM. katae
Ranomafana: Beremby, Sahadikaina-21.246147.5222644betsileanusM. katae
Ranomafana: Beremby, Sahalavabe-21.207547.5313860betsileanusM. katae
stelligerM. stelliger
Ranomafana: Beremby, Sahalavakely-21.210947.5304780stelligerM. stelliger
Ranomafana: Bibiango (Old Bridge)-21.257847.41831106betsileanusM. katae
Ranomafana: Fompohonina III-21.265147.42251027*biporusM. glosi
betsileanusM. katae
Ranomafana: Imaloka-21.242747.46511052tricinctusM. grubenmanni
Ranomafana: Kidonavo bridge-21.226247.36961152betsileanusM. betsileanus
betsileanusM. katae
Ranomafana: Maharira-21.325847.40251248betsileanusM. betsileanus
betsileanusM. katae
biporusM. bletzae
Ranomafana: near Park entrance-21.257347.4211936*betsileanusM. betsileanus
Ranomafana: Ranomafana National Park, Ampangadiamesa-21.242347.41091147betsileanusM. katae
Ranomafana: Ranomafana-21.250647.41781132betsileanusM. betsileanus
National Park, AndranovoromaintybetsileanusM. katae
Ranomafana: Ranomafana,-21.261447.4413710betsileanusM. betsileanus
AmbodirianabetsileanusM. katae
Ranomafana: Ranomafana, Antenna-21.260947.445795betsileanusM. betsileanus
Ranomafana: Ranomafana, Imaloka-21.242147.46521020betsileanusM. katae
tricinctusM. grubenmanni
Ranomafana: Ranomafanakely-21.248747.37181134betsileanusM. betsileanus
curtusM. alutus
Ranomafana: Sahateza, Pond Donald-21.257947.35971182betsileanusM. katae
Ranomafana: Sakaroa-21.264847.41221046*betsileanusM. katae
Ranomafana: Talatakely, Piste E-21.262847.4257959betsileanusM. katae
biporusM. glosi
Ranomafana: Valohoaka-21.298647.43861091*betsileanusM. betsileanus
betsileanusM. katae
Ranomafana: Vatoharanana-21.289047.42941000betsileanusM. katae
Ranomafana: Vohiparara,-21.235247.39611170betsileanusM. betsileanus
SahamalaotrabetsileanusM. katae
road from Ranomafana to Tolongoina-21.353647.6078468betsileanusM. betsileanus
Sahafina-Brickaville-18.810348.980064*betsileanusM. incognitus
biporusM. eulenbergeri
fergusoniM. georgei
Sahamalaza-14.248847.9553170ulcerosusM. ulcerosus
Sahambaky Forest (Lakato)-19.06548.34980betsileanusM. betsileanus
Sahavontsira-16.905749.2217466biporusM. brevirostris
inaudaxM. manerana antsanga
tricinctusM. grubenmanni
Sainte Luce-24.766747.183330tricinctusM. gudrunae
Sainte Luce: forest at QMM climate station-24.779847.171323tricinctusM. gudrunae
Sambava region-14.274650.166412*fergusoniM. fergusoni
Sampanandrano (Anosy Mountains)-24.139947.0742539betsileanusM. katae
Site 1 near Ambodimandresy-13.713349.4911778ulcerosusM. steinfartzi
small stream next to Sofia, along road-15.70748.5807252ulcerosusM. ulcerosus
Sorata: above campsite-13.683149.4411398– 1417inaudaxM. manerana fotaka
Sorata: above campsite, bamboo-13.674649.44061516betsileanusM. jonasi
forestinaudaxM. manerana fotaka
Sorata: above campsite, cloud forest-13.675849.4381599inaudaxM. manerana fotaka
Sorata: Camp-13.685149.44171279inaudaxM. manerana fotaka
Sorata: gallery forest at creek near-13.722149.4385776betsileanusM. jonasi
Andrafainkona, AmbararataulcerosusM. bellyi
Sorata (outside forest)-13.713949.4966844*ulcerosusM. bellyi
Tampolo forest (Analanjirofo)-17.288749.41167fergusoniM. georgei
Toamasina-18.166749.383320fergusoniM. georgei
Tolagnaro-25.033346.9560betsileanusM. tripunctatus
Torotorofotsy-18.874748.3725960betsileanusM. betsileanus
betsileanusM. katae
Torotorofotsy: Camp Prolemur-18.769248.4266956betsileanusM. betsileanus
Tsaranoro: Forêt Sacrée-22.081246.7747940betsileanusM. betsileanus
curtusM. bourgati
curtusM. mahery
Tsaratanàna-13.9848.84NAbetsileanusM. jonasi
ulcerosusM. ulcerosus
Tsaratanàna: Camp 0 (Ankijagna Lagnana)-14.239948.97211162betsileanusM. jonasi
Tsaratanàna: Manarikoba forest, Andampy-14.042248.7617730ulcerosusM. schulzi
Tsaratanàna: Manarikoba forest, Antsahamanara-14.04548.78531000ulcerosusM. steinfartzi
Tsingy de Bemaraha: Camp 1, Antranopasazy-18.708644.7189120curtusM. mahery
Tsingy de Bemaraha: Camp 2, Andafiabe on Beboka River-18.784244.7794177curtusM. mahery
Tsingy de Bemaraha: Camp 3, Bendrao Forest-18.784444.8603427curtusM. mahery
Tsingy de Bemaraha: Camp 4, Bendrao Forest-18.797244.8814420curtusM. mahery
Tsinjoarivo-19.6847.7711610betsileanusM. betsileanus
curtusM. alutus
Tsinjoarivo: Camp 3, Vatateza-19.719947.8571319betsileanusM. betsileanus
Tsitongambarika: Andranomaizina-24.583847.147442betsileanusM. tripunctatus
tricinctusM. gudrunae
Tsitongambarika: Ivohibe-24.561247.1924415betsileanusM. tripunctatus
tricinctusM. gudrunae
Vatolampy-20.82847.3191580curtusM. curtus
Vohidrazana-18.966148.5097810betsileanusM. incognitus
betsileanusM. katae
biporusM. eulenbergeri
way to Sampanandrano-24.137247.0949411*betsileanusM. katae