The discovery of two new species in the Cyrtodactylus irregularis group highlights that hidden diversity remains in the largest clade of the mega-diverse genus Cyrtodactylus Author Ngo, Hanh Thi 6B87B552-B68B-46A7-8678-4E6EAE01428A Institute of Zoology, University of Cologne, Zülpicher Str. 47 b, 50674 Cologne, Germany. & Cologne Zoo, Riehler Str. 173, D- 50735 Cologne, Germany. & Central Institute for Natural Resources and Environmental Studies, Vietnam National University, Hanoi, 19 Le Thanh Tong Street, Hanoi, Vietnam. ngohanhhus@gmail.com Author Hormann, Helene 0BBD64A8-84F4-4ED0-BD64-18703C3CBB26 Institute of Zoology, University of Cologne, Zülpicher Str. 47 b, 50674 Cologne, Germany. & Cologne Zoo, Riehler Str. 173, D- 50735 Cologne, Germany. helene099@arcor.de Author Le, Minh Duc 785D605E-6A6F-4A67-BD18-7F40F426FC2C Central Institute for Natural Resources and Environmental Studies, Vietnam National University, Hanoi, 19 Le Thanh Tong Street, Hanoi, Vietnam. & Faculty of Environmental Sciences, Hanoi University of Science, Vietnam National University, Hanoi, 334 Nguyen Trai Road, Hanoi, Vietnam. & Department of Herpetology, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024, USA. le.duc.minh@hus.edu.vn Author Pham, Cuong The 24C187A9-8D67-4D0E-A171-1885A25B62D7 Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi 10072, Vietnam. & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Cau Giay, Hanoi 10072, Vietnam. cuongiebr@gmail.com Author Phung, Trung My F05DF764-A42C-4993-B570-04E351E31A7D Dong Khoi 9 A, Tam Hiep, Bien Hoa, Dong Nai Province, Vietnam. pmytrung@gmail.com Author Do, Dang Trong 81C099A4-09B1-4156-BAC4-B8532C3A03E8 Phu Yen University, 01 Nguyen Van Huyen Road, Tuy Hoa City, Phu Yen Province, Vietnam. dotrongdang@gmail.com Author Ostrowski, Sabrina 30862A2D-E20E-4FC2-B81C-A7E5FBF3E1C7 Institute of Zoology, University of Cologne, Zülpicher Str. 47 b, 50674 Cologne, Germany. & Cologne Zoo, Riehler Str. 173, D- 50735 Cologne, Germany. sabrinaostrowski@mail.de Author Nguyen, Truong Quang F080FC36-C4C5-45CB-BFDB-E0598662A4DD Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi 10072, Vietnam. & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Cau Giay, Hanoi 10072, Vietnam. nqt2@yahoo.com Author Ziegler, Thomas 5716DB92-5FF8-4776-ACC5-BF6FA8C2E1BB Institute of Zoology, University of Cologne, Zülpicher Str. 47 b, 50674 Cologne, Germany. & Cologne Zoo, Riehler Str. 173, D- 50735 Cologne, Germany. ziegler@koelnerzoo.de text European Journal of Taxonomy 2023 2023-06-19 875 1 70 100 http://dx.doi.org/10.5852/ejt.2023.875.2141 journal article 10.5852/ejt.2023.875.2141 1ad43c61-d534-4bd0-ae9f-bc907ed8744c 2118-9773 8064396 CC306FFC-76D4-4D8F-8E87-01E92594483A
Table 4 (continued on next page).Morphological comparisons between Cyrtodactylus chumuensis sp. nov., Cyrtodactylus arndti sp. nov. and 23 congeners from the Cyrtodactylus irregularis complex based on examination of specimens and data obtained from the literature (Ziegler et al. 2002; Nguyen et al. 2006, 2013, 2014, 2015, 2017a, 2017b, 2021; Heidrich et al. 2007; Orlov et al. 2007; Nazarov et al. 2008, 2012; Ngo 2088, 2013; Ngo & Bauer 2008; Ngo et al. 2010; Rösler et al. 2008; Ngo & Chan 2010, 2011; Ngo & Grismer 2012; Ziegler et al. 2010, 2013; Luu et al. 2011, 2017; Phung et al. 2014; Le et al. 2016; Ostrowski et al. 2020, 2021; Do et al. 2021). Abbreviations:? = data not available in literature; – = characteristic not present; + = characteristic present but not uniquely determined; * = regenerated or broken tail.
SpeciesSVL (mm)TaL (mm)VEFSFPPP (M)PP (F)LD4LT4Color pattern of dorsumSubcaudals
1Cyrtodactylus chumuensis sp. nov.67.551.4*43–454–50–26–7?16–1917–21bandedabsent
2Cyrtodactylus arndti sp. nov.73.4–80.950.1*–91.5126–385–110–26615–2017–22bandedpresent
3C. badenensis59.3–74.158.6–82.425–29absentabsent00?18–22bandedpresent
4C. bidoupimontis74.0–86.375.0–8638–438–10absent4–6015–2018–23bandedabsent
5C. bugiamapensis58.6–76.865.3–83.036–466–10absent7–110–715–1717–20blotchedabsent
6C. buchardi60.0–65.046.0–54.030absentabsent901412blotchedabsent
7C. caovansungi90.4–94.0120.038–4486902223–25bandedpresent
8C. cattienensis43.5–69.051–64.728–423–8absent6–80–8 pitted scales12–1614–19bandedabsent
9C. chungi66.6–68.562.8*–82.230–314–6absent76?17–20bandedabsent
10C. cryptus62.5–90.863.5–88.447–50absentabsent9–11018–1920–23bandedabsent
11C. cucdongensis55.8–65.9max. 81.341–445–9absent5–64–613–1815–20bandedabsent
12C. culaochamensis69.8–79.889.7–91.245–50absentabsent7–8absent18–1920–23bandedabsent
13C. datimax 70.1max 57.342–484–73–4 each5–6??18–19blotchedabsent
14C. gialaiensis50.1–62.8?38–45absentabsent9–100–814–1515–17bandedabsent
15C. huynhi67.2–79.861.5–78.643–463–53–87–90–8 pitted scales14–1717–21bandedabsent
16C. irregularis72.0–86.066.0–74.038–457–8absent5–70–615–1618–19blotchedabsent
17C. kingsadai83.0–94.0max 11739–469–120–47–94–819–2121–25bandedpresent
18C. orlovi61–77.7Max 71.236–393–8absent5–6015–1716–19bandedabsent
19C. phnomchiensis76.1–80.756.9–79.145–540–8absent4–51–7 pitted scales18–2020–23bandedabsent
20C. phumyensis63.6–66.8?33–415–7absent5–76 pitted scales18–1918–21bandedabsent
21C. phuocbinhensis46.0–60.476.143–475absent7016–2117–19striped/blotchedabsent
22C. pseudoquadrivirgatus48.6–83.355.7–82.341–57absentabsent5–95–1015–2116–25blotchedabsent
23C. raglai95–111.7113.4–13536–399–10absent50?21–22bandedpresent
24C. sangi49.9–56.347.9*374absent74 pitted scale??bandedabsent
25C. takouensis74.7–81.177.7–91.039–403–50–23–4016–1718–20bandedpresent
26C. taynguyenensis60.0–85.066.0–94.042–49absentabsent6013–1817–21blotchedabsent
27C. yangbayensis78.5–92.391.3–109.139–465–160–26–8016–1915–17bandedpresent
28C. ziegleri84.6–93.095.0–107.033–398–100–65–80–816–1918–21bandedabsent