| T. otavalo sp. nov. | T. ucucha | T. igor sp. nov. | T. silvestris |
| Tail very long (LT about 144 % of HB) | Tail long (LT about 128 % of HB) | Tail long (LT about 130 % of HB) | Tail long (LT about 140 % of HB) |
| White tail tip (15–35 mm) | Tail tip usually not white (when present <5 mm) | Tail tip usually not white (when present <5 mm) | Tail tip usually not white (when present <5 mm) |
| Dorsal pelage uniformly grayish brown | Dorsal pelage uniformly dark grayish brown | Dorsal pelage verona Brown, darker in the midline | Dorsal pelage uniformly dark grayish brown |
| Incipient rostral tube present | Rostral tube absent | Rostral tube absent | Rostral tube absent |
| Thomas angle 80 ° | Thomas angle 86 ° | Thomas angle 80 ° | Thomas angle 80 ° |
| Zygomatic arches converge anteriorly | Widely flaring zygomatic arches | Zygomatic arches converge anteriorly | Zygomatic arches converge anteriorly |
| Upper first molar aligned with posterior edge of the zygomatic plate | First upper molar does not reach the posterior edge of the zygomatic plate | First upper molar does not reach the posterior edge of the zygomatic plate | Upper first molar reaches beyond the posterior edge of the zygomatic plate |
| Incisive foramina long (LIF about 63 % of LD), usually not extending posteriorly between molar alveoli | Incisive foramina vey short (LIF about 57 % of LD), not approaching level of molar alveoli | Incisive foramina long (LIF about 72 % of LD), usually not extending posteriorly between molar alveoli | Incisive foramina long (LIF about 65 % of LD), usually not extending posteriorly between molar alveoli |
| Upper edge of the mastoid does not exceed the edge of the subsquamosal fenestra | The superior border of the mastoid goes above the subsquamosal fenestra. | Upper edge of the mastoid does not exceed the edge of the subsquamosal fenestra | Upper edge of the mastoid does not exceed the edge of the subsquamosal fenestra |
| Capsular process of lower incisor alveolus usually indistinct | Capsular process of lower incisor alveolus usually distinct | Capsular process of lower incisor alveolus indistinct or absent | Capsular process of lower incisor alveolus indistinct or absent |
| Upper incisors opisthodont, not procumbent | Upper incisors orthodont, conspicuously procumbent | Upper incisors opisthodont, not procumbent | Upper incisors opisthodont, not procumbent |
| Subsquamosal fenestra larger than the postglenoid foramen | Subsquamosal fenestra usually smaller than postglenoid foramen | Subsquamosal fenestra larger than the postglenoid foramen | Subsquamosal fenestra larger than the postglenoid foramen |
| M 3 Paraflexo short and metaflexo indistinct | Paraflexus and metaflexus short | Paraflexus and metaflexus length and fused | Paraflexus and mesoflexus short |
| M 3 comparatively large | M 3 comparatively small | M 3 comparatively large | M 3 comparatively small |
| Procingulum of m 1 divided | Procingulum of m 1 undivided | Procingulum of m 1 divided | Procingulum of m 1 undivided or indistinct |