Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam Author Bernardes, Marta Cologne Zoo, Riehler Str. 173, 50735 Cologne, Germany & Terrestrial Ecology, Institute of Zoology, University of Cologne, Zuelpicher Str. 47 b, 50674 Cologne, Germany mrtbernardes@gmail.com Author Le, Minh Duc Faculty of Environmental Sciences, University of Science, Vietnam National University, Hanoi, 334 Nguyen Trai Road, Hanoi, Vietnam & Central Institute for Natural Resources and Environmental Studies, Vietnam National University, Hanoi, 19 Le Thanh Tong, Hanoi, Vietnam & Department of Herpetology, American Museum of Natural History, Central Park West at 79 th Street, New York, New York 10024, USA Author Nguyen, Truong Quang Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam https://orcid.org/0000-0002-6601-0880 Author Pham, Cuong The Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam Author Pham, Anh Van Faculty of Natural Science and Technology, Tay Bac University, Quyet Tam Ward, Son La City, Son La Province, Vietnam Author Nguyen, Tao Thien Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam & Vietnam National Museum of Nature, 18 Hoang Quoc Viet St., Hanoi, Vietnam Author Roedder, Dennis Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, 53113 Bonn, Germany Author Bonkowski, Michael Terrestrial Ecology, Institute of Zoology, University of Cologne, Zuelpicher Str. 47 b, 50674 Cologne, Germany Author Ziegler, Thomas Cologne Zoo, Riehler Str. 173, 50735 Cologne, Germany & Terrestrial Ecology, Institute of Zoology, University of Cologne, Zuelpicher Str. 47 b, 50674 Cologne, Germany text ZooKeys 2020 935 121 164 http://dx.doi.org/10.3897/zookeys.935.37138 journal article http://dx.doi.org/10.3897/zookeys.935.37138 1313-2970-935-121 4D41FCF4A59544F095EFCF38D0FC127C 33263B29FD6E5F15BCFC61C058095F9C

Table 5. Results from the MANOVA of the log-transformed ratio of characters to SVL of males. The variation was analyzed between the populations from Son La and Thanh Hoa provinces (taxon 1; N = 9) and the populations from Phu Tho and Hoa Binh provinces (taxon 2; N = 10), and between these (jointly referred to as Tylototriton cf. asperrimus) and the population from Lai Chau Province (taxon 3; in the comparison based on head and dorsum related data: NT. cf. asperrimus = 19 and N taxon3 = 3; and in the comparison of limb related data, NT. cf. asperrimus = 21 and N taxon3 = 4). F: F -test; DF: degrees of freedom; P: p -value. For abbreviations of characters see Materials and methods. In Bold significant results.

	taxon 1 x taxon 2			T. cf. asperrimus x taxon 3
	F	DF	P	F	DF	P
MHW	17.62	17	<0.001	0.08	20	0.79
HW	7.48	17	0.01	52.48	20	<0.001
EN	4.85	17	0.04	7.52	20	0.01
HL	0.76	17	0.4	0.09	20	0.77
IN	4.56	17	0.05	0.09	20	0.77
LJL	2.51	17	0.13	2.22	20	0.15
RAD/HUM	5.92	17	0.03	0.34	23	0.56
FEM	21.13	17	<0.001	1.10	23	0.30
TIB/FEM	7.07	17	0.02	0.42	23	0.52
HIND	7.47	17	0.01	0.43	23	0.52
HIND/FORE	4.95	17	0.04	0.03	23	0.87
HUM	0.14	17	0.72	4.66	23	0.04
RAD	0.17	17	0.7	<0.001	23	0.98
TIB	1.99	17	0.18	0.83	23	0.78
FORE	0.01	17	0.91	1.87	23	0.19
TL	1.10	17	0.31	0.07	20	0.79
TH	0.35	17	0.56	0.71	20	0.41
TL.TH	0.25	17	0.63	0.56	20	0.46
L5N	0.84	17	0.37	12.43	20	<0.01
WVr	1.61	17	0.22	5.02	20	0.04
TkL	0.90	17	0.36	3.74	20	0.07