Johnius sasakii, a new species of croaker (Perciformes: Sciaenidae) with a key to Johnius from East Malaysia, Borneo Author Hanafi, Norhafiz Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan Author Chen, Meng-Hsien Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan & Marine Ecology and Conservation Research Center, National Academy of Marine Research, Kaohsiung, Taiwan & Institute of Marine Ecology and Conservation, National Sun Yat-sen University, Kaohsiung, Taiwan Author Seah, Ying Giat Faculty of Fisheries and Food Science, University Malaysia Terengganu (UMT), Kuala Nerus, Malaysia & Fish Division, South China Sea Repository and Reference Centre, Institute of Oceanography and Environment, University Malaysia Terengganu (UMT), Kuala Nerus, Malaysia Author Chang, Chih-Wei Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan & Marine Ecology and Conservation Research Center, National Academy of Marine Research, Kaohsiung, Taiwan & National Museum of Marine Biology and Aquarium, and Graduate Institute of Marine Biology, National Dong Hwa University, Pingtung, Taiwan Author Liu, Shang Yin Vanson 0000-0002-9472-3941 Department of Marine Biotechnology and Resources, National Sun Yat-Sen University, Kaohsiung, Taiwan oceandiver 6426 @ gmail. com; https: // orcid. org / 0000 - 0002 - 9472 - 3941 oceandiver6426@gmail.com Author Chao, Ning Labbish National Museum of Marine Biology and Aquarium, and Graduate Institute of Marine Biology, National Dong Hwa University, Pingtung, Taiwan & Bio-Amazonia Conservation International, Boston, Massachusetts, USA. text Zootaxa 2022 2022-03-21 5116 3 393 409 journal article 20163 10.11646/zootaxa.5116.3.5 7d00b0d3-235f-4460-a1a2-d1ac95dd75f3 1175-5326 6372220 E04B56E0-5B56-4413-B8D6-552DDCEEB111

TABLE 2. Morphometric and meristic data of type specimen of Johnius sasakii sp. nov. in present study. SD = standard deviation.

	Holotype	Paratypes (17 individuals)
TABLE 2. (Continued)
*Abbreviation: D1 means to first dorsal-fin spine length; P2 means to pelvic-fin length; and P1 means to pectoral-fin length.
	NMMB-P 34733	Min	Max	Mean	SD
Standard Length (mm)	110.9	100.0	146.9	117.7
Head Length (mm)	34.4	29.2	44.2	35.6
Eye Diameter (mm)	7.4	6.6	8.8	7.6
Counts
1st dorsal-fin spine	10	10	10	10	0.0
2nd dorsal-fin spine	1	1	1	1	0.0
2nd dorsal-fin soft rays	27	26	31	28	0.0
Anal-fin spine	2	2	2	2	0.0
Anal-fin soft rays	7	7	7	7	0.0
Pectoral-fin rays	17	11	18	16	0.0
Outer gill rakers of 1st arch	15	13	15	14	0.0
upper limb	6	4	6	5	0.0
lower limb	9	8	10	9	0.0
Inner gill rakers of 1st arch	11	10	13	11	0.0
upper limb	3	3	4	3	0.0
lower limb	8	6	9	8	0.0
Pored lateral line scales	51	37	49	47	0.0
Scales above lateral line	6	4	6	5	0.0
Scales below lateral line	9	6	14	10	0.0
Circumpeduncular scales	14	11	17	14	0.0
Vertebrae	23	23	23	23	0.0
precaudal	10	10	10	10	0.0
caudal	13	13	13	13	0.0
Measurements as %HL
Snout length	23.2	23.0	29.5	26.3	2.0
Maxillary length	41.3	34.0	42.6	38.2	2.8
Eye diameter	21.5	18.0	24.4	21.6	2.0
Interorbital width	23.4	23.8	27.9	26.0	1.3
2nd spine length	31.6	24.9	35.6	29.2	2.6
1st ray length	39.6	30.7	46.7	36.3	3.5
	NMMB-P 34733	Min	Max	Mean	SD
Measurements as %SL
Snout to anal-fin origin	69.3	67.1	73.3	70.6	1.5
Snout to 2nd dorsal-fin origin	53.2	49.3	55.0	51.7	1.7
Snout to 1st dorsal-fin origin	33.3	31.8	36.2	33.7	1.3
Snout to pectoral-fin insertion	31.2	27.4	32.8	30.3	1.5
Snout to pelvic-fin insertion	35.4	31.0	36.5	34.0	1.5
Pectoral-fin length	21.3	16.6	24.9	19.7	2.3
Head length	31.0	27.7	32.3	30.2	1.2
Pelvic-fin length	15.3	12.7	20.0	16.1	2.0
Body depth (D1-P2)	27.0	25.2	29.0	27.1	1.2
Body width (P1-P1)	15.6	13.4	18.5	15.5	1.8
1st dorsal-fin base length	20.1	15.9	23.1	19.3	2.0
longest spine length	14.9	11.9	16.7	13.8	1.3
2nd dorsal-fin base length	40.1	35.8	44.8	39.7	2.5
longest ray length	8.9	7.0	14.4	9.8	1.7
Anal-fin base length	9.5	6.8	10.2	8.8	0.9
2nd spine length	9.8	7.4	10.2	8.5	0.8
1st ray length	12.3	9.0	13.3	10.9	1.0
Caudal-peduncle depth	10.6	8.7	10.7	9.5	0.4
Pelvic-fin tip to vent	12.7	13.8	22.8	18.4	2.7
Vent to anal-fin origin	4.4	3.2	7.9	5.7	1.2
Measurements as %ED
Longest gill raker on 1st arch	14.9	11.3	19.9	14.6	2.4
Longest gill filament on 1st arch	55.7	37.7	59.9	51.9	3.4