Johnius sasakii, a new species of croaker (Perciformes: Sciaenidae) with a key to Johnius from East Malaysia, Borneo Author Hanafi, Norhafiz Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan Author Chen, Meng-Hsien Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan & Marine Ecology and Conservation Research Center, National Academy of Marine Research, Kaohsiung, Taiwan & Institute of Marine Ecology and Conservation, National Sun Yat-sen University, Kaohsiung, Taiwan Author Seah, Ying Giat Faculty of Fisheries and Food Science, University Malaysia Terengganu (UMT), Kuala Nerus, Malaysia & Fish Division, South China Sea Repository and Reference Centre, Institute of Oceanography and Environment, University Malaysia Terengganu (UMT), Kuala Nerus, Malaysia Author Chang, Chih-Wei Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan & Marine Ecology and Conservation Research Center, National Academy of Marine Research, Kaohsiung, Taiwan & National Museum of Marine Biology and Aquarium, and Graduate Institute of Marine Biology, National Dong Hwa University, Pingtung, Taiwan Author Liu, Shang Yin Vanson 0000-0002-9472-3941 Department of Marine Biotechnology and Resources, National Sun Yat-Sen University, Kaohsiung, Taiwan oceandiver 6426 @ gmail. com; https: // orcid. org / 0000 - 0002 - 9472 - 3941 oceandiver6426@gmail.com Author Chao, Ning Labbish National Museum of Marine Biology and Aquarium, and Graduate Institute of Marine Biology, National Dong Hwa University, Pingtung, Taiwan & Bio-Amazonia Conservation International, Boston, Massachusetts, USA. text Zootaxa 2022 2022-03-21 5116 3 393 409 journal article 20163 10.11646/zootaxa.5116.3.5 7d00b0d3-235f-4460-a1a2-d1ac95dd75f3 1175-5326 6372220 E04B56E0-5B56-4413-B8D6-552DDCEEB111

TABLE 3. Frequency distribution of selected meristics and morphometrics of Johnius sasakii sp. nov., J. carouna and J. heterolepis from the South China Sea region.

		Upper limb of outer				Lower limb of outer gill rakers						Total outer gill rakers of 1st arch
		gill rakers
Counts	n	4	5	6	Mean	8	9	10	11	12	Mean	13	14	15	16	17	18	Mean
Johnius sasakii	18	3	11	4	5.1	2	14	2			9.0	5	7	6				14.1
sp. nov. Johnius carouna	11		5	6	5.5			1	6	4	11.3				3	7	1	16.8
Johniusheterolepis	11	10	1		4.1		1	9	1		10.0		10	1				14.1