Johnius sasakii, a new species of croaker (Perciformes: Sciaenidae) with a key to Johnius from East Malaysia, Borneo Author Hanafi, Norhafiz Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan Author Chen, Meng-Hsien Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan & Marine Ecology and Conservation Research Center, National Academy of Marine Research, Kaohsiung, Taiwan & Institute of Marine Ecology and Conservation, National Sun Yat-sen University, Kaohsiung, Taiwan Author Seah, Ying Giat Faculty of Fisheries and Food Science, University Malaysia Terengganu (UMT), Kuala Nerus, Malaysia & Fish Division, South China Sea Repository and Reference Centre, Institute of Oceanography and Environment, University Malaysia Terengganu (UMT), Kuala Nerus, Malaysia Author Chang, Chih-Wei Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan & Marine Ecology and Conservation Research Center, National Academy of Marine Research, Kaohsiung, Taiwan & National Museum of Marine Biology and Aquarium, and Graduate Institute of Marine Biology, National Dong Hwa University, Pingtung, Taiwan Author Liu, Shang Yin Vanson 0000-0002-9472-3941 Department of Marine Biotechnology and Resources, National Sun Yat-Sen University, Kaohsiung, Taiwan oceandiver 6426 @ gmail. com; https: // orcid. org / 0000 - 0002 - 9472 - 3941 oceandiver6426@gmail.com Author Chao, Ning Labbish National Museum of Marine Biology and Aquarium, and Graduate Institute of Marine Biology, National Dong Hwa University, Pingtung, Taiwan & Bio-Amazonia Conservation International, Boston, Massachusetts, USA. text Zootaxa 2022 2022-03-21 5116 3 393 409 journal article 20163 10.11646/zootaxa.5116.3.5 7d00b0d3-235f-4460-a1a2-d1ac95dd75f3 1175-5326 6372220 E04B56E0-5B56-4413-B8D6-552DDCEEB111

TABLE 4. Comparison of select meristic and morphometric characters among congeneric species of Johnius (Johnius) from East Malaysia

	J. sasakii sp. nov.	J. carouna	J. heterolepis	J. macrorhynus
*Parentheses indicate as mean and standard deviation.
	(n=18)	(n=11)	(n=11)	(n=10)
Voucher number	NMMB-P34733-34739	NMMB-P34740-34743	NMMB-P34748-34751	NMMB-P34744-34747
Type status	Paratype	Voucher specimens	Voucher specimens	Voucher specimens
Type locality	East Malaysia	East Malaysia	East Malaysia	East Malaysia
Standard length (mm)	100.0–146.9	87.8–141.4	88.1–112.2	116.5–188.3
Counts
Dorsal-fins rays	26–31	28–34	25–28	27–28
Scales above lateral line	4–6	5–7	4–5	4–5
Scales below lateral line	6–14	8–11	8–10	7–9
Number of lower limb of outer gill rakers	8–10	10–12	9–11	7–8
Total outer gill rakers of 1st arch	13–15	16–18	14–15	11–13
Measurement (%SL)
Head length	27.7–32.3(30.2±1.2)	25.9–30.7(27.7±1.2)	30.4–32.7(31.4±0.8)	28.8–31.4(30.4±0.7)
Body depth	25.2–29.0(27.1±1.2)	23.2–28.0(25.0±1.4)	23.8–27.9(26.1±1.3)	26.0–32.2(27.7±1.7)
2nd spine of anal-fin length	7.4–10.2(8.8±0.8)	11.3–13.8(12.3±0.8)	8.7–11.0(9.6±0.6)	6.3–8.5(7.5±0.7)
1st ray of anal-fin length	9.0–13.3(10.9±1.0)	10.9–14.7(13.4±1.1)	10.9–13.9(12.5±0.8)	9.8–12.0(10.9±0.7)
Anal-fin base length	6.8–10.2(8.8±0.9)	9.0–10.6(9.8±0.5)	9.7–10.8(10.1±0.4)	8.2–11.0(9.5±0.9)
Pre-anal length	67.1–73.6(70.6±1.5)	65.9–70.3(68.0±1.4)	67.2–74.7(70.1±2.4)	66.9–72.2(70.1±1.8)
Measurement (%HL)
Snout length	23.0–29.5(26.3±2.0)	23.8–28.6(26.3±1.4)	21.1–29.4(25.6±2.5)	27.5–32.3(29.9±1.2)
Upper jaw length	34.0–42.6(38.3±2.8)	38.0–43.8(41.0±1.8)	35.0–42.8(40.4±2.6)	35.5–43.5(40.7±2.3)
Tip upper jaw to hinge length	26.4–35.4(29.1±2.7)	24.4–29.7(26.5±1.5)	27.6–33.0(29.9±1.7)	24.2–29.8(26.8±1.6)
Tip lower jaw to hinge length	21.2–29.0(24.9±2.5)	21.8–26.3(24.0±1.4)	24.7–29.8(26.8±1.4)	19.7–24.8(22.3±1.5)
Eye diameter	18.0–24.4(21.6±2.0)	20.9–26.6(23.3±1.6)	19.3–24.7(21.2±1.5)	19.0–24.4(21.9±1.5)
Interorbital width	23.8–27.9(26.0±1.3)	22.0–29.3(26.6±2.3)	20.3–28.2(24.9±2.6)	21.6–25.7-23.8±1.4)
2nd spine of anal-fin length	24.9–35.6(29.2±2.6)	38.8–49.4(44.5±2.7)	27.4–34.1(30.7±1.8)	16.8–27.8(22.8±2.8)
1st ray of anal-fin length	30.7–46.7(36.3±2.8)	43.2–54.3(48.6±2.5)	35.2–44.6(39.9±2.5)	30.2–37.3(33.8±2.5)
Measurement (%ED)
Gill raker length	11.8–19.9(14.6±2.4)	13.6–26.9(15.8±2.8)	11.7–19.0(16.6±2.4)	4.8–11.1(9.7±0.7)