Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae) Author Dayrat, Benoit Department of Biology, Pennsylvania State University, University Park, PA 16802, USA https://orcid.org/0000-0002-1514-4854 bdayrat@gmail.com Author Goulding, Tricia C. Department of Biology, Pennsylvania State University, University Park, PA 16802, USA Author Apte, Deepak Bombay Natural History Society, Hornbill House, Opp. Lion Gate, Shaheed Bhagat Singh Road, Mumbai 400 001, Maharashtra, India Author Aslam, Sadar Centre of Excellence in Marine Biology, University of Karachi, Karachi 75270, Pakistan https://orcid.org/0000-0002-4340-7885 Author Bourke, Adam College of Engineering, Information Technology and the Environment, Charles Darwin University, Ellengowan Dr, Casuarina, NT 0810, Australia Author Comendador, Joseph National Museum of the Philippines, Taft Ave, Ermita, Manila, 1000, Metro Manila, Philippines Author Khalil, Munawar Department of Marine Science, Universitas Malikussaleh, Reuleut Main Campus, Kecamatan Muara Batu, North Aceh, Aceh, 24355, Indonesia https://orcid.org/0000-0002-8264-5317 Author Ngo, Xuan Qu ảng Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam https://orcid.org/0000-0003-2587-1999 Author Tan, Siong Kiat Lee Kong Chian Natural History Museum, 2 Conservatory Dr, National University of Singapore, 117377, Singapore Author Tan, Shau Hwai Centre for Marine and Coastal Studies, Universiti Sains Malaysia, 11800, Minden Penang, Malaysia & Marine Science Laboratory, School of Biological Sciences, Universiti Sains Malaysia, 11800, Minden Penang, Malaysia text ZooKeys 2020 972 1 224 http://dx.doi.org/10.3897/zookeys.972.52853 journal article http://dx.doi.org/10.3897/zookeys.972.52853 1313-2970-972-1 791674942E9242C38D1FD4DE7264D7B7 2751774FF66A5579AE66145BE5055C8E

Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions.

Species		Max. animal length (mm)	Papillae with dorsal eyes	Intestinal type (transitional loop orientation)	Retractor muscle insertion (near heart or end of visceral cavity)	Muscular sac length (mm)	Accessory penial gland spine length (mm)	Accessory penial gland spine diameter base (μm)	Accessory penial gland spine diameter tip (μm)	Penis hooks length (μm)	Distribution
P. peronii		140	15-20	I (12-3 o'clock)	heart (exc. vestigial)	>20	3.0-5.0	400-500	160-200	<50	Indo-West Pacific (Zanzibar to Okinawa and Tonga)
P. okinawensis		27	10-15	I (12-3 o'clock)	heart	<15	1.8-2.3	240-300	115-150	<35	Japan (Okinawa)
P. madagascariensis		80	12-18	V	end of VC	<15	2-2.4	200-230	70-80	<100	Western Indian Ocean
P. platei		30	7-10	V	end of VC	<5	0.7-1.0	65-100	20-30	<60	Western Pacific (PNG to Hawaii & French Polynesia)
P. setoensis		20	8-12	V	end of VC	<5	0.9-1.2	80-85	15-25	<25	Japan (Wakayama)
P. griffithsi		25	6-10	V	end of VC (exc. heart)	<5	0.50-0.62	60-65	15-20	<25	Indo-West Pacific (Mauritius to Kei Islands & New Ireland)
P. sydneyensis		50	8-16	I (3-6 o'clock)	end of VC	<10	0.6-1.0	90-100	20-50	<30	Queensland, New South Wales & New Caledonia
P. willani		65	10-25	I (3-6 o'clock)	end of VC	<25	1.5-1.9	240-250	80-100	<37	Northern Territory
P. verruculata	#1	60 (exc. 73)	10-22	I (3-6 o'clock)	end of VC	<15	1.4-2.0	100-270	35-50	<50	Singapore to eastern Australia, New Caledonia & Japan
	#2	55	14-22			<10	1.4-1.7	140-160	30-35		Sumatra & Andaman
	#3	40	10-18				1.8-2.2	200-270	40-80	<60	Malaysia & Singapore
	#4	60					(1.3) 2.2-2.8	200	50		Western India, Pakistan & Persian Gulf
	#5	50	10-20				1.8-2.0	150-180	45-50	<55	Mozambique & Madagascar
	#6	40	10-18			<15	2.0-2.4	140-200	55-60		Red Sea