Description of a new species of Southeast Asian reed snake from northern Laos (Squamata: Colubridae: Genus Calamaria F. Boie, 1827) with a revised diagnosis of Calamaria yunnanensis Chernov, 1962 Author Lee, Justin L. text Journal of Natural History 2021 2021-06-14 55 9 - 10 531 560 http://dx.doi.org/10.1080/00222933.2021.1909165 journal article 3258 10.1080/00222933.2021.1909165 f5778971-d9e4-4877-b9c3-fa5fbf31bfdf 1464-5262 5476206 AD8D2EAF-E76E-4D49-9DA8-4C5A656B4F9A

Table 3. Morphological comparisons between Calamaria nebulosa sp. nov. and specimens of Calamaria yunnanensis. Data with asterisks are adapted from Zhao et al. (1998) and Yang and Rao (2008). Abbreviations are noted in the materials and methods, except for DSR6R, which stands for the number of dorsal scales reducing to six rows above position of subcaudal anterior to tail tip; DSR4R, which stands for the number of dorsal scales reducing to four rows above position of subcaudal anterior to tail tip; and ‘SC’, which stands for subcaudal. Entries marked with ‘ – ’ are not available.
Morphological character	nebulosa sp. nov.	yunnanensis	yunnanensis	yunnanensis	yunnanensis	yunnanensis	yunnanensis
Museum catalogue	FMNH 258666	KIZ 054176	ZISP 17073	ROM 41547	KIZ 056009	KIZ 056010	KIZ 056011
Locality	Phongsaly Province, Laos	Jingdong, Yunnan Province, China	Jingdong, Yunnan Province, China	Simao, Yunnan Province, China	Jingdong, Yunnan Province, China	Jingdong, Yunnan Province, China	Jingdong, Yunnan Province, China
Sex	F	F	M	M	M	M	Unknown
SVL	326	490	225	226	280*	280	136
TailL	28	26	20	20	20	16	8
TotalL	354	516	245	246	300*	296	144
TailL/totalL ratio	0.079	0.050	0.082	0.081	0.067*	0.054	0.059
HeadL	8.8	9.8	–	7.4	–	7.6	4.8
HeadW	5.6	6.8	–	4.0	–	4.9	3.3
EyeD	1.3	0.8	–	1.0	–	0.6	0.6
SnL	2.8	3.3	–	2.6	–	2.5	2.0
SnW	2.1	2.8	–	2.0	–	1.7	1.2
NarEye	2.0	2.5	–	1.5	–	1.2	1.3
IOD	3.5	4.0	–	3.1	–	2.1	2.0
Dorsal scales	13–13–13	13–13–13	13–13–13	13–13–13	13–13–13	13–13–13	13–13–13
Gulars (preventrals)	3	3	–	3	–	3	3
Ventrals	179	199	173	167	184*	179	181
Cloacal plate	Single	Single	Single	Single	Single	Single	Single
Subcaudals	22	19	20	19	20	15	16
Total body scales	202	219	194	187	205*	195	198
Subcaudal ratio	0.109	0.087	0.103	0.102	0.098	0.073	0.081
Maxillary teeth	9/9	9*	–	8/8	–	9*	9*
Supralabials	4/4	4/4	4	4/4	4/4*	4/4	4/4
Infralabials	5/5	5/5	5	5/5	6/6*	5/5	5/5
Preocular	0/0	0/0	0	0/0	0/0	0/0	0/0
Postocular	1/1	1/1	1	1/1	1/1	1/1	1/1
Scales around paraparietal DSR6R	6/6 12th SC	6/6	6 –	6/6 3rd SC	6/6 3rd SC	6/6 5th SC	6/6 –
DSR6R	12th SC	2nd SC	–	6/6 3rd SC	6/6 3rd SC	6/6 5th SC	–
DSR4R	Last SC	3rd to last SC	–	3rd to last SC	3rd to last SC	3nd to last SC	–
End of tail	Obtuse point	Obtuse point	–	Obtuse point	Obtuse point	Obtuse point	Obtuse point
Dorsal colour pattern	Five longitudinal stripes	Longitudinal stripes barely visible	Longitudinal stripes barely visible	Longitudinal stripes across flanks	Longitudinal stripes barely visible	Longitudinal stripes barely visible	Longitudinal stripes across flanks
Margins of ventral scales	Immaculate	Immaculate	–	Edged with black	Edged with black	Edged with black	Edged with black
Nuchal collar	Absent	Absent	Present	Present	Present	Present	Present
Midventral stripe on tail	Absent	Absent	Present	Present	Present	Present	Present
Source	This study	This study; Yang and Rao (2008)	Chernov (1962)	This study	This study; Yang and Rao (2008)	This study; Yang and Rao (2008)	This study; Yang and Rao (2008)