Description of a new species of Southeast Asian reed snake from northern Laos (Squamata: Colubridae: Genus Calamaria F. Boie, 1827) with a revised diagnosis of Calamaria yunnanensis Chernov, 1962 Author Lee, Justin L. text Journal of Natural History 2021 2021-06-14 55 9 - 10 531 560 http://dx.doi.org/10.1080/00222933.2021.1909165 journal article 3258 10.1080/00222933.2021.1909165 f5778971-d9e4-4877-b9c3-fa5fbf31bfdf 1464-5262 5476206 AD8D2EAF-E76E-4D49-9DA8-4C5A656B4F9A

Table 4. Morphological comparisons between Calamaria nebulosa sp. nov. (FMNH 258666), Calamaria yunnanensis and all species of Calamaria known from mainland Southeast Asia (see Materials and methods for list of literature sources). Symbols used in this table include: (M) = male; (F) = female; (m) = modified maxillary teeth.

Morphological character	C. nebulosa sp. nov.	C. abramovi	C. albiventer	C. andersoni	C. buchi	C. concolor	C. dominici	C. gialaiensis	C. ingeri	C. lovii gimletii
(Continued)
Portion of rostral scale visible from above greater than half with length of prefrontal suture (1) or less than or equal to to half its length (0)	0	–	1	1	1	–	0	0	0	0
Number of scales surrounding paraparietal	6	6	5	6	5	5	6	5	6	6
Prefrontal: longer than frontal scale (1) or smaller than/equal to frontal scale (0)	0	0	0	0	0	1	0	0	1	0
Preocular: present (1) or absent (0)	0	1	1	1	1	1	0	1	1	0
Mental touching (1) or separated from (0) anterior genials	0	0	1	0	1	1	1	1	0	0
Posterior genials meeting in midline (1), diverging or meeting only in anteriorly (0)	0	0	1	0	1	1	0	0	1	1
End of tail	Obtuse point	Sharp point	Sharp point	Obtuse point	Obtuse point	Obtuse point	Obtuse point	Round	Blunt	Blunt
Dorsal scales reducing to four rows above position of subcaudal anterior to tail tip	Last subcaudal	Last subcaudal	5th–8th subcaudal	2nd-to-last subcaudal	3rd–4th subcaudal	Last subcaudal	5th–6th subcaudal	Last subcaudal	Last subcaudal	Last–5th subcaudal
Maxillary teeth (modified = m)	9 (m)	8 (m)	9 (m)	9 (m)	9 (m)	8 (m)	9 (m)	9 (m)	7 (m)	8–9 (m)
Ventrals	179 (F)	174 (F) 159 (M)	147–162 (F) 143–144 (M)	171 (M)	221–236 (F)	3 + 209 (M)	174 (F)	3 + 191 (M)	213–228 (F)	215–249 (F) 161–202 (M)
Subcaudals	22 (F)	20 (F) 26 (M)	15–19 (F) 21–22 (M)	23 (M)	13–14 (F)	19 (M)	17–18 (F)	23 (M)	11–10 (F)	10–12 (F) 14–20 (M)
Dorsum	Bluish-grey	Black	Brown	Brownish	Black	Brown	Black	Grey-brown	Dark brown	Dark brown
Venter	Yellow	Black and yellow	Red	Yellow	Yellow	Cream	Yellow and black	Yellow	Yellow	Yellow
Dorsal scales with light spots present (1) or absent (0)	1	1	1	1	1	0	0	0	0	0
Dorsum with light nuchal collar (1) or absent (0)	0	0	1	0	0	0	0	0	1	0–1
Tail with rings: present (1) or absent (0)	0	0	0	0	0	0	0	1	1	0–1
Underside of tail with dark midventral stripe present (1) or absent (0)	0	0	1	1	0	1	0	1	0	0
TailL/TotalL ratio (given as %)	7.9 (F)	7.1 (F) 13.3 (M)	4.7–8.8 (F) 8.8–9.3 (M)	9.2 (M)	3.9–4.1 (F)	7.3 (M)	6.0 (F)	8.8 (M)	3.5–4.2 (F)	3.0–3.7 (F) 6.2–8.4 (M)
Morphological character C. strigiventris	C. lovii ingermarxorum	C. lumbricoidea C. thanhi	C. pavimentata C. yunnanensis	C. sangi	C. schlegeli schlegeli	C.				septentrionalis
Portion of rostral scale visible from above greater than half with length of prefrontal suture (1) or less than or equal to to half its length (0)	1	0	1	1	0	0	1		1	0
Number of scales surrounding paraparietal	6	4–5	5–6	5–6	5–6	6	6		6–7	6
Prefrontal: longer than frontal scale (1) or smaller than/equal to frontal scale (0)	1	0	0	0	0	0	0		0	0
Preocular: present (1) or absent (0)	0	1	1	1	0–1	1	1		0	0
Mental touching (1) or separated from (0) anterior genials	1	1	0	1	0	0	0		0	0
Posterior genials meeting in midline (1), diverging or meeting only in anteriorly (0)	1	1	1	0	0–1	1	0		0	0
End of tail	Blunt	Sharp point	Sharp point	Obtuse point	Blunt	Round	Abruptly to point		Gradually to point	Obtuse point
Dorsal scales reducing to four rows above position of subcaudal anterior to tail tip	4th–5th subcaudal	Last–11th subcaudal	Last–13th subcaudal	Last–3rd subcaudal	3rd–25th subcaudal	Does not decrease	Last–6th subcaudal		Does not decrease	3rd-to-last subcaudal
Maxillary teeth (modified = m)	8 (m)	9–11 (m)	8–9 (m)	9 (m)	9–10 (m)	8–9 (m)	9–11 (m)		9 (m)	8–9 (m)
Ventrals	205 (M)	137–229 (F) 144–196 (M)	137–206 (F) 125–168 (M)	2 + 190 (?)	136–180 (F) 129–161 (M)	168–188 (F) 148–166 (M)	158–180 (F) 130–157 (M)		198 (F) 184 (M)	199 (F) 167–184 (M)
Subcaudals	23 (M)	13–21 (F) 17–27 (M)	8–20 (F) 13–33 (M)	19 (?)	19–37 (F) 25–44 (M)	6–11 (F) 15–19 (M)	20–30 (F) 29–33 (M)		21 (F) 28 (M)	19 (F) 15–20 (M)
Dorsum	Bluish-grey	Dark brown to black	Brown	Greyish brown	Dark brown to black	Dark brown to black	Slate grey to grey-brown		Dark blue to grey	Blue-grey to olive-brown
Venter	Dark grey	Yellow	Yellow	Yellow	white or yellow	Yellow	Yellow and black		Yellow	Yellow or red
Dorsal scales with light spots present (1) or absent (0)	0	0–1	0	0	0	1	0		0	1
Dorsum with light nuchal collar (1) or absent (0)	1	0	0–1	1	0	1	0		0	0–1
Tail with light rings: present (1) or absent (0)	0	0–1	0–1	0	0–1	1	0		1	0
Underside of tail with dark midventral stripe present (1) or absent (0)	0	0	0–1	1	0	1	1		0	0–1
TailL/TotalL ratio (given as %)	7.4 (M)	3.9–8.3 (F) 6.3–11.4 (M)	3.7–8.5 (F) 6.9–16.9 (M)	6.2 (?)	7.3–14.4 (F) 11.1–21.3 (M)	2.6–4.3 (F) 6.3–8.6 (M)	8.4–11.5 (F) 13.8–17.9 (M)		6.8 (F) 9.9 (M)	5.0 (F) 5.4–8.2 (M)