Carlgren’s hesitation allayed: redescription and systematics of Heteranthus verruculatus Klunzinger, 1877 (Cnidaria, Actiniaria), with a redefinition of Heteranthidae Carlgren, 1900 Author Yap, Nicholas Wei Liang Author Quek, Zheng Bin Randolph Author Tan, Ria Author Nugroho, Dharma Arif Author Lee, Jen Nie Author Lee Berumen, Michael Author Tan, Koh Siang Author Huang, Danwei text Contributions to Zoology 2021 2020-11-27 90 2 155 182 http://dx.doi.org/10.1163/18759866-bja10015 journal article 268186 10.1163/18759866-BJA10015 ef0a5d8c-febb-411a-ba02-ba847d325c93 1875-9866 8343299

TABLE 2 Cnidom of Heteranthus verruculatus Klunzinger, 1877, comprising type specimens (i.e,. those of H.verruculatus and H.insignis), and voucher specimens. Regarding the latter, cnidae measurements were taken from both solitary and clustering polyps

		Type Specimens					Voucher Specimens
		NRS 4861		NRS 4076		Solitary polyps			Clustering polyps
Tissue	Cnidae	Range length n		Range length × n		Range length × N		n	Range length ×	N	n
		× width		width		width			width
Marginal	Spirocysts (A)	(10.0)12.0–	10	(9.6)10.8–18.0	10	(11.0)13.0–23.0	6/6	60	10.0–20.2 ×	7/7	70
tentacles		17.0 × 1.5–3.0		× 2.4		× 2.0–3.0			(1.5)2.0–3.0
	Basitrichs (B)	(14.0)16.5–	10	15.6–18.0 ×	10	14.0–18.7 ×	6/6	60	(12.1)13.0–19.0 ×	7/7	70
		19.0 × 2.0–3.0		2.4–3.0(3.6)		2.0–3.0			2.0–3.0
		14.8–18.3 ×	?
		2.0–2.5
Discal tentaclesSpirocysts (C)		‒	‒	13.2–16.8 ×	10	(8.0)10.0–19.0	6/6	60	(9.0)11.1–17.2 ×	5/5	50
				2.4–3.0		× 2.0–3.5			2.0–3.0(3.5)
	Basitrichs (D)	‒	‒	10.8–13.2 × 2.4	10	9.0–16.0 ×	6/6	60	8.0–20.5(22.0) ×	5/5	50
						2.0–3.0			(1.5)2.0–2.5(3.0)
Marginal	Basitrichs (E)	8.0–10.0 ×	10	12.0–18.6(24.0) × 5		10.0–15.0 ×	6/6	59	7.5–13.1 ×	7/7	71
projections		1.5–2.0		2.4–3.0(3.6)		2.0–2.5			2.0–2.5
Column	Basitrichs (F)	18.0–21.0 ×	10	(12.0)15.0–25.2 × 23		11.0–19.2 ×	6/6	60	11.0–17.0 ×	7/7	80
		2.0–3.0		2.0–3.0		2.0–3.0(3.5)			(1.5)2.0–2.5
		15.5–20.6 ×	?
		2.0–2.5
Actino-pharynxBasitrichs (G)		‒	‒	(18.6)20.4–23.4 0 10		(15.0)16.0–25.3 6/6		60	16.0–23.5 ×	7/7	70
				× 2.4–3.0		× 2.0–4.0			2.0–3.0(3.5)
	Microbasic p- masti-	‒	‒	15.6–18.6 ×	10	15.0–25.3 ×	6/6	60	15.0–25.5 ×	7/7	71
	gophores (H)			(2.4)3.6–4.8		3.0–6.0			3.0–7.0