Verbascum birjandense and V. urumiense (Scrophulariaceae), two new endemic species for Flora of Iran Author Sotoodeh, Arash 0000-0003-3735-5481 UMR Agroécologie, Université de Bourgogne, France & Central Herbarium of Tehran University, School of Biology, College of Science, University of Tehran, Iran & EDB, UMR 5174, Université Toulouse III Paul Sabatier, 118 route de Narbonne, F- 31062 Toulouse cedex 9, France laure. civeyrel @ univ-tlse 3. fr; https: // orcid. org / 0000 - 0003 - 3735 - 5481 Author Attar, Farideh Central Herbarium of Tehran University, School of Biology, College of Science, University of Tehran, Iran Author Laitung, Beryl UMR Agroécologie, Université de Bourgogne, France Author Civeyrel, Laure 0000-0003-3735-5481 EDB, UMR 5174, Université Toulouse III Paul Sabatier, 118 route de Narbonne, F- 31062 Toulouse cedex 9, France laure. civeyrel @ univ-tlse 3. fr; https: // orcid. org / 0000 - 0003 - 3735 - 5481 text Phytotaxa 2022 2022-03-03 538 1 35 44 journal article 20297 10.11646/phytotaxa.538.1.3 96605e3b-6f83-42e4-8b34-1d053c40b4b3 1179-3163 6331962

TABLE 2. Morphological comparison between Verbascum erianthum, V. songaricum, and V. urumiense, based on Flora Iranica and new observations. The diagnostic characters of the first are highlighted in bold.

	V. erianthum	V. songaricum	V. urumiense
Plant height	0.6–2 m	0.3–1.5 m	1.7–1.9 m
Indumentum	greyish, or yellowish stellate hairs all along the plant	greyish, or whitish stellate hairs along the plant	whitish stellate hairs all along the plant
Stem	rounded, leafy, above ± branched	rounded or striated, and obscurely angular, with sparse or numerous branches	angular, leafy
Basal leaves	oblong or ovate-oblong, 10-30 × 5-12 cm, obtuse denticulate or angular-crenate, rarely subentire	oblong- to linear-lanceolate, 10–40 × 1.5–12 cm, ± distinctly crenulated or entire, acutish to acuminate	elliptic, 30–41 × 8–12 cm, entire, broadly acute.
Stem leaves	upper stem leaves not undulated	upper stem leaves not or a little undulated	upper stem leaves not undulated
Petiole of basal leaves	3 to 10 cm, unwinged	2–7 cm, winged	absent
Pedicels	(2–)4–10 mm long	5–12 mm long	4–6 mm long
Pedicel/calyx ratio	2 times longer	equal or a little longer	equal or a little longer
Calyx	4–6 mm long	6–10 mm long	3–5 mm long
Corolla	yellow, 15–30 mm diam., without pellucid glands. outside stellate hairy, inside glabrous or sparsely ciliated	yellow, 25–35 mm diam., without pellucid glands, outside stellate hairy, glabrous inside	yellow, 25–30 mm diam., with pellucid glands, outside stellate hairy, inside sparse ciliated
Stamen	5, filaments with orange hairs	5, filaments with whitish-yellow hairs	5, filaments with whitish hairs
Anthers	reniform	all reniform, or 3 reniform and the 2 inferior decurrent	reniform
Capsule	capsule ellipsoid-ovoid to pyramidal-ovoid, 3.5–6.5 × 3–5 mm, covered by stellate hairs to glabrous	ovoid to ovoid-ellipsoid, 5–7 × 4–5 mm, tomentose, glabrescent	not seen