The Stoor Hobbit of Guangdong: Goniurosaurus gollum sp. nov., a cave-dwelling Leopard Gecko (Squamata, Eublepharidae) from South China Author Qi, Shuo State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China Author Wang, Jian State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China https://orcid.org/0000-0003-4249-7767 Author Grismer, L. Lee Herpetology Laboratory, Department of Biology, La Sierra University, Riverside, California 92515, USA Author Chen, Hong-Hui State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China Author Lyu, Zhi-Tong State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China https://orcid.org/0000-0002-4891-3859 Author Wang, Ying-Yong State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China wangyy@mail.sysu.edu.cn text ZooKeys 2020 991 137 153 http://dx.doi.org/10.3897/zookeys.991.54935 journal article http://dx.doi.org/10.3897/zookeys.991.54935 1313-2970-991-137 2D9EEFC0B43E4AC386E789944E54169B A789ECB84D295459B43890139564D4C4

Table 4. Diagnostic characters distinguishing Goniurosaurus gollum sp. nov. from all other known species of Goniurosaurus. Data come from Grismer et al. 1999, 2002; Orlov et al. 2008; Ziegler et al. 2008; Blair et al. 2009; Wang et al. 2010, 2013, 2014; Yang and Chan 2015; Zhou et al. 2018; Qi et al. 2020; Zhu et al. 2020.

Character	G. kuroiwae group	G. lichtenfelderi group	G. luii group	G. yingdeensis group (4 species)
	(5 spp.)	(5 spp.)	(7 spp.)	Goniurosaurus gollum sp. nov.	G. varius	G. yingdeensis	G. zhelongi
Scales of upper eyelid one-half the size of scales on the top of head or equal in size	Equal	Equal	Equal or 1/2	Equal	Equal	Equal	Equal
Enlarged row of supraorbital tubercles	Absent	Absent or present	Absent or present	Absent	Absent	Absent or indistinct	Absent or indistinct
Eyelid fringe scales	<52	43-77	41-67	59-63	50-56	46-64	42-53
No. of paravertebral tubercles	Unknown	23-36	27-38	25-26	27-29	22-33	28-33
Dorsal tubercle rows at midbody	Unknown	19-22	20-25	16-17	21-24	20-25	23-28
Scales around midbody	Unknown	95-140	112-147	121-128	101-110	101-116	99-109
Nasal scales surrounding nares	Unknown	8-9	5-9	7-8	7-9	7-11	6-8
Internasals	Unknown	1	0-3	1	1-2	1-3	1-2
Tubercles between orbits	Present or absent	Present or absent	Present or absent	Absent	Present	Present	Absent
Claws sheathed by scales	Absent	Present	Present	Present	Present	Present	Present
Lateral scales of claw sheaths	Absent	Long, curved	Long, curved	Short, shell-shaped	Short, shell-shaped	Short, shell-shaped	Short, shell-shaped
No. of precloacal pores in males	0	17-46	16-33	10-11	10	10-13	9-12
Posterior shape of nuchal loop	Rounded	Protracted or rounded	Protracted	Rounded	Rounded	Rounded	Rounded
No. of body bands between nuchal loop and the caudal constriction	3 or 4	3 or 4	3, 4 or 5	2,3 or 4	4	4	4
Dark spotting in body bands	Present or absent	Present or absent	Present or absent	Absent	Present or absent	Absent	Absent
Lateral spotting on belly present or absent	Absent	Absent	Present or absent	Present	Present	Present	Present