Paraharmotrema karinganiense n. gen., n. sp. (Digenea: Liolopidae) infecting the intestine of serrated hinged terrapin (Pelusios sinuatus), east African black mud turtle (Pelusios subniger), and South African helmeted turtle (Pelomedusa galeata) and a phylogenetic hypothesis for liolopid genera Author Dutton, Haley R. Aquatic Parasitology Laboratory and Southeastern Cooperative Fish Parasite and Disease Laboratory, School of Fisheries, Aquaculture, & Aquatic Sciences, College of Agriculture, Auburn University, 559 Devall Dr., Auburn, AL, 36832, USA hzd0026@auburn.edu Author DuPreez, Louis H. African Amphibian Conservation Research Group, Unit for Environmental Sciences and Management, North-West University, Private Bag × 6001, Potchefstroom, 2520, South Africa & South African Institute for Aquatic Biodiversity, Somerset Street, Grahamstown 6139, South Africa Author Urabe, Misako University Shiga Prefecture, Department of Ecosystem Studies, Faculty of Environmental Science, 2500 Hassaka, Hikone, Shiga, 5228533, Japan Author Bullard, Stephen A. Aquatic Parasitology Laboratory and Southeastern Cooperative Fish Parasite and Disease Laboratory, School of Fisheries, Aquaculture, & Aquatic Sciences, College of Agriculture, Auburn University, 559 Devall Dr., Auburn, AL, 36832, USA text International Journal for Parasitology: Parasites and Wildlife 2022 2022-04-30 17 43 52 http://dx.doi.org/10.1016/j.ijppaw.2021.09.006 journal article 292466 10.1016/j.ijppaw.2021.09.006 04e04e2c-6c95-4c7a-ad1b-18f2050dcef0 2213-2244 PMC8688888 34976724 10668222

Table 2 Key to accepted genera of Liolopidae.

1a.	Testes in anterior half of body	Helicotrema Odhner, 1912
1b.	Testes in posterior half of body	2
2a.	Cirrus sac abutting ventral sucker	Liolope Cohn, 1902
2b.	Cirrus sac not abutting ventral sucker	3
3a.	Testes near, in posterior 1/3 of body	Dracovermis Brooks and
		Overstreet, 1978
3b.	Testes far apart, not limited to posterior	4
	1/3 of body
4a.	Testes smooth, not deeply lobed	Harmotrema Nicoll, 1914
4b.	Testes deeply lobed	Paraharmotrema Dutton & Bullard
		n. gen.