Phylogenetic and multivariate analyses of Gekko smithii Gray, 1842 recover a new species from Peninsular Malaysia and support the resurrection of G. albomaculatus (Giebel, 1861) from Sumatra Author Grismer, L. Lee Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92515, USA lgrismer@lasierra.edu Author del Pinto, Lelani Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92515, USA Author Quah, Evan S. H. https://orcid.org/0000-0002-5357-1953 Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400, Kota Kinabalu, Sabah, Malaysia & School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia Author Anuar, Shahrul School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia & Center for Marine and Coastal Studies, Universiti Sains Malaysia, 11800 USM, Pulau Pinang, Malaysia Author Cota, Michael Natural History Museum, National Science Museum, Thailand Author McGuire, Jimmy A. Museum of Vertebrate Zoology and Department of Integrative Biology, University of California, Berkeley, California 94720, USA Author Iskandar, Djoko T. The Indonesian Academy of Sciences, Jakarta 10110 Indonesia Author Wood Jr, Perry L. https://orcid.org/0000-0003-3767-5274 Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan 48109 - 1085, USA Author Grismer, Jesse L. https://orcid.org/0000-0002-2542-5430 Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92515, USA text Vertebrate Zoology 2022 2022-02-03 72 47 80 http://dx.doi.org/10.3897/vz.72.e77702 journal article http://dx.doi.org/10.3897/vz.72.e77702 2625-8498-72-47 3A5B7AD25A8142E9B0136A95ED03E7B6 66A14C58EB6659F29033A6E475A606B4

Table 9. Summary statistics of the meristic data. sd = 1 standard deviation. N = sample size. Character abbreviations are in the Materials and methods.

Gekko smithii	SL	IL	IN	FS	CS	MB	PVT	LRT	VS	TL1	TL4	PP
mean (+/- sd)	13.7 (+/- 1.48)	11.6 (+/- 1.28)	4.5 (+/- 0.75)	22.6 (+/- 2.06)	6.7 (+/- 1.03)	112.9 (+/- 9.73)	19.1 (+/- 1.94)	9.7 (+/- 0.61)	29.2 (+/- 3.17)	17.6 (+/- 1.39)	21.4 (+/- 1.5)	13.2 (+/- 1.95)
range	11-17	9-14	3-6	19-26	4-9	94-137	17-23	8-11	23-35	15-20	19-24	13-15
N	27	27	27	27	27	27	27	27	27	27	27	23
Gekko hulk sp. nov.
mean (+/- sd)	13.3 (+/- 0.94)	11 (+/- 0.92)	4.2 (+/- 0.55)	19.5 (+/- 2.1)	6 (+/- 0.91)	98.7 (+/- 6.82)	18.4 (+/- 1.31)	10.1 (+/- 0.57)	26.0 (3.74)	16.6 (+/- 1.27)	20.7 (+/- 1.23)	8.7 (+/- 1.85)
range	10-15	9-13	3-5	13-24	4-8	84-110	16-21	9-11	22-28	14-19	18-24	6-13
N	42	42	42	42	42	42	42	42	42	42	42	16
cf. Gekko albomaculatus
mean (+/- sd)	14 (+/- 1.00)	11.4 (+/- 1.27)	4.7 (+/- 0.49)	21.4 (+/- 2.7)	7.4 (+/- 0.53)	101 (+/- 5.69)	18.7 (+/- 1.8)	10.3 (+/- 0.76)	30.1 (+/- 2.41)	16.3 (+/- 1.5)	19.7 (+/- 1.5)	14.0 (+/- 3.00)
range	13-16	10-14	4-5	17-26	7-8	90-109	16-21	9-11	26-33	14-18	17-21	11-17
N	7	7	7	7	7	7	7	7	7	7	7	3
cf. Gekko albofasciolatus
mean (+/- sd)	14 (+/- 0.82)	11.6 (+/- 0.98	5.3 (+/- 0.49)	21.9 (+/- 1.77)	6.7 (+/- 0.49)	98.7 (+/- 3.35)	18.3 (+/- 1.5)	10.4 (+/- 0.98)	27.9 (+/- 4.1)	15.7 (+/- 1.8)	19.4 (+/- 1.99)	13.6 (+/- 0.96)
range	13-15	11-13	5 or 6	19-24	6 or 7	94-103	16-20	9-12	23-33	14-19	17-22	13-15
N	7	7	7	7	7	7	7	7	7	7	7	4