Integrative taxonomy supports the description of two new species of Macrobiotidae (Tardigrada, Eutardigrada) from Kristianstads Vattenrike Biosphere Reserve in Sweden Author Hulterström, Jens Department of Zoology, Swedish Museum of Natural History, Box 50007, 104 05 Stockholm, Sweden. jens.hulterstrom15@gmail.com Author Guidetti, Roberto Department of Life Sciences, University of Modena and Reggio Emilia, 41124 Modena, Italy. roberto.guidetti@unimore.it Author Jönsson, K. Ingemar Department of Environmental Science, Kristianstad University, SE- 291 88 Kristianstad, Sweden. ingemar.jonsson@hkr.se Author Atherton, Sarah Department of Zoology, Swedish Museum of Natural History, Box 50007, 104 05 Stockholm, Sweden. sarah.atherton@nrm.se text European Journal of Taxonomy 2025 2025-12-08 1030 1 52 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3135/13991 journal article 313536 10.5852/ejt.2025.1030.3135 0e837015-e9de-4bbe-92b1-4fbb915238a4 2118-9773 17962351 60D8486F-0A30-423A-832B-3B5BF58C20ED

Appendix 1 (continued on next 14 pages). GenBank Accession numbers for COI, 18S, ITS, and 28S sequences of all specimens analysed in this study. New sequences are highlighted in red.

Taxon	Specimen	COI	18S	ITS	28S	Taxon name in GenBank	Notes	Reference in GenBank
Diaforobiotus islandicus	IS.042	MT808072	MT812470	MT812597	MT812461	Diaforobiotus islandicus		Stec et al. 2020d
Macrobiotus aff. polonicus	S165.1	MW593929	MW588026	MW588021	MW588032	Macrobiotus aff. polonicus S165		Vecchi & Stec 2021
Macrobiotus aff. polonicus	S165.2	MW593930	MW588027	MW588020	MW588033	Macrobiotus aff. polonicus S165		Vecchi & Stec 2021
Macrobiotus almadai	ZYL123	MW990245	MW995180			Macrobiotus almadai		Wang et al. (unpubl.)
Macrobiotus annewintersae	S207.1	MW593927	MW588025	MW588019	MW588030	Macrobiotus annewintersae		Vecchi & Stec 2021
Macrobiotus annewintersae	S207.2	MW593928	MW588024	MW588018	MW588031	Macrobiotus annewintersae		Vecchi & Stec 2021
Macrobiotus ariekammensis ariekammensis	NO.393.01	MZ460999	MZ463668	MZ463657	MZ463674	Macrobiotus ariekammensis ariekammensis		Stec et al. 2022
Macrobiotus ariekammensis groenlandicus	GL.018.04	MZ461006	MZ463664	MZ463654	MZ463679	Macrobiotus ariekammensis groenlandicus		Stec et al. 2022
Macrobiotus azzunae	C4218_T5	MW698698				Macrobiotus sp. 1 MC-2021	temporary taxon name has not been updated in GenBank at time of writing	Marnissi et al. 2021
Macrobiotus azzunae	C4218_V4	MW698697	MW695447	MW695454	MW695450	Macrobiotus sp. 1 MC-2021	temporary taxon name has not been updated in GenBank at time of writing	Marnissi et al. 2021
Macrobiotus basiatus	USA /NEL/1	MT502116	MT498094	MT505165	MT488397	Macrobiotus sp. 1 DG-2020	temporary taxon name has not been updated in GenBank at time of writing	Nelson et al. 2020
Macrobiotus birendrai	CN8.101	MW656266	MW680641	MW680418	MW680644	Macrobiotus birendrai		Kayastha et al. 2021
Macrobiotus caelestis	KG.007	MK737922	MK737073	MK737072	MK737071	Macrobiotus caelestis		Coughlan et al. 2019
Macrobiotus canaricus	ES.004_H1	MH057765	MH063925	MH063928	MH063934	Macrobiotus canaricus		Stec et al. 2018c
Macrobiotus canaricus	ES.004_H2	MH057766				Macrobiotus canaricus		Stec et al. 2018c
Macrobiotus cf. hufelandi	C2959a	HQ876590		OP596303		Macrobiotus cf. hufelandi 1 MC-2011		Bertolani et al. 2011
Macrobiotus cf. muralis	C3251_2	OP561785	OP596299			Macrobiotus cf. muralis		Bertolani et al. 2023
Macrobiotus cf. muralis	C3251_3	OP561786	OP596300			Macrobiotus cf. muralis		Bertolani et al. 2023
Macrobiotus cf. muralis	C3251_4	OP561788				Macrobiotus cf. muralis		Bertolani et al. 2023
Macrobiotus cf. muralis	C3251_5	OP561789				Macrobiotus cf. muralis		Bertolani et al. 2023
Macrobiotus cf. muralis	C3251_FA	OP561787	OP596301			Macrobiotus cf. muralis		Bertolani et al. 2023
Macrobiotus cf. nelsonae	1		HQ604965			Macrobiotus		Bertolani et al. 2014
Macrobiotus cf. recens	ES.006 haplotype 1	MH057768	MH063927	MH063932	MH063936	Macrobiotus cf. recens DS-2018		Stec et al. 2018c
Macrobiotus cf. sapiens	S12.Mac.1	OK662997	OK663226	OK663215	OK663237	Macrobiotus cf. sapiens		Vecchi et al. 2022c
Macrobiotus cf. shonaicus	MM1-2018	LC431582		LC431591		Macrobiotus cf. shonaicus		Matsumoto & Sugiura (unpubl.)
Macrobiotus crustulus	GF.271	MT260371	MT261912	MT261907	MT261903	Macrobiotus crustulus		Stec et al. 2020a
Macrobiotus dolosus	C3209_2	OP561772	OP596290			Macrobiotus dolosus		Bertolani et al. 2023
Macrobiotus dolosus	C3209_4	OP561774				Macrobiotus dolosus		Bertolani et al. 2023
Macrobiotus dolosus	C3209_ US 2	OP561773				Macrobiotus dolosus		Bertolani et al. 2023
Macrobiotus dolosus	C3581_V6	OP561775	OP596292			Macrobiotus dolosus		Bertolani et al. 2023
Macrobiotus engbergi	haplotype 1	MN444824	MN443039	MN443036	MN443034	Macrobiotus engbergi		Vecchi et al. 2024b
Macrobiotus engbergi	haplotype 2	MN444825		MN443037		Macrobiotus engbergi		Vecchi et al. 2024b
Macrobiotus fontourai	C2861_5	OP561783	OP596295			Macrobiotus fontourai		Bertolani et al. 2023
Macrobiotus fontourai	C2861_ US 1	OP561784	OP596296			Macrobiotus fontourai		Bertolani et al. 2023
Macrobiotus glebkai	UA.003	MW246134	MW247177	MW247180	MW247176	Macrobiotus glebkai		Kiosya et al. 2021
Macrobiotus hannae	PL.010	MH057764	MH063922	MH063923	MH063924	Macrobiotus hannae		Nowak & Stec 2018
Macrobiotus hufelandi	C2953a	HQ876585	OP596302			Macrobiotus hufelandi		Bertolani et al. 2023
Macrobiotus hufelandi	S605.Mac.1	OK662992	OK663221	OK663210	OK663232	Macrobiotus hufelandi		Bertolani et al. 2023
Macrobiotus joannae	1		HQ604974			Macrobiotus joannae		Bertolani et al. 2014
Macrobiotus kamilae	IN.030 haplotype 1	MK737920	MK737070	MK737067	MK737064	Macrobiotus kamilae		Coughlan & Stec 2019
Macrobiotus kathyae	US.006.01	PP386934	PP391285	PP391289	PP391287	Macrobiotus sp. MV2024	temporary taxon name has not been updated in GenBank at time of writing	Massa & Vecchi 2024
Macrobiotus kathyae	US.006.02	PP386935	PP391286	PP391290	PP391288	Macrobiotus sp. MV2024	temporary taxon name has not been updated in GenBank at time of writing	Massa & Vecchi 2024
Macrobiotus kathyae	US.006.03	PP386932				Macrobiotus sp. MV2024	temporary taxon name has not been updated in GenBank at time of writing	Massa & Vecchi 2024
Macrobiotus kathyae	US.006.04	PP386933				Macrobiotus sp. MV2024	temporary taxon name has not been updated in GenBank at time of writing	Massa & Vecchi 2024
Macrobiotus kirghizicus	KG.062.01	MZ461002	MZ463666	MZ463659	MZ463672	Macrobiotus kirghizicus		Stec et al. 2022
Macrobiotus kosmali	M8.1	OP141639	OP142472	OP153786	OP143765	Macrobiotus kosmali		Kayastha et al. 2023a
Macrobiotus kristenseni	C3291_A02_V2	KC193573	KC193577			Macrobiotus kristenseni		Guidetti et al. 2013
Macrobiotus kyoukenus	C4313_ US 3	ON809462	ON818314	ON818301	ON818309	Macrobiotus kyoukenus		Cesari et al. 2022
Macrobiotus macrocalix	PL.110	MH057767	MH063926	MH063931	MH063935	Macrobiotus macrocalix		Stec et al. 2018c
Macrobiotus margoae	US.057	MN888315	MN888368	MN888340	MN888354	Macrobiotus margoae		Vecchi & Stec 2021
Macrobiotus margoae	US.057.1	MT807927	MT809072	MT809098	MT809084	Macrobiotus margoae		Stec et al. 2021b
Macrobiotus margoae	US.057.3	MT807928		MT809099		Macrobiotus margoae		Stec et al. 2021b
Macrobiotus mileri	IL.001.01	OR544397	OR543312	OR543316	OR543320	Macrobiotus mileri		Stec 2024
Macrobiotus mileri	IL.001.02	OR544398	OR543313	OR543317	OR543321	Macrobiotus mileri		Stec 2024
Macrobiotus mileri	IL.001.03	OR544399				Macrobiotus mileri		Stec 2024
Macrobiotus mileri	IL.001.04	OR544400				Macrobiotus mileri		Stec 2024
Macrobiotus noongaris	AU.031	MK737919	MK737069	MK737066	MK737063	Macrobiotus noongaris		Coughlan & Stec 2019
Macrobiotus ovovittatus	GL.001.01	OR544395	OR543311	OR543314	OR543318	Macrobiotus ovovittatus		Stec 2024
Macrobiotus pallarii	IT.337.1	MT807924	MT809069	MT809094	MT809081	Macrobiotus pallarii		Stec et al. 2021b
Macrobiotus pallarii	IT.337.2	MT807925	MT809070	MT809095	MT809082	Macrobiotus pallarii		Stec et al. 2021b
Macrobiotus pallarii	IT.337.3	MT807926	MT809071	MT809096	MT809083	Macrobiotus pallarii		Stec et al. 2021b
Macrobiotus papei	TZ.027	MH057763	MH063881	MH063921	MH063880	Macrobiotus papei		Stec et al. 2018b
Macrobiotus paulinae		KT 951668	KT935502	KT935500		Macrobiotus paulinae		Stec et al. 2015
Macrobiotus scoticus	DK.056.1	OK662989	OK663217	OK663206	OK663228	Macrobiotus scoticus		Vecchi et al. 2022c
Macrobiotus shonaicus	haplotype 1	MG757136	MG757132	MG757134	MG757133	Macrobiotus shonaicus		Stec et al. 2018a
Macrobiotus siderophilus	C2796_2	OP561776	OP596293			Macrobiotus siderophilus		Bertolani et al. 2023
Macrobiotus siderophilus	C3282_UFK	OP561777				Macrobiotus siderophilus		Bertolani et al. 2023
Macrobiotus siderophilus	C3282_UFL	OP561778				Macrobiotus siderophilus		Bertolani et al. 2023
Macrobiotus siderophilus	C3282_UFM	OP561779				Macrobiotus siderophilus		Bertolani et al. 2023
Macrobiotus siderophilus	C3282_UFN	OP561780	OP596294			Macrobiotus siderophilus		Bertolani et al. 2023
Macrobiotus siderophilus	C3282_UFO	OP561781				Macrobiotus siderophilus		Bertolani et al. 2023
Macrobiotus siderophilus	C3282_UFP	OP561782				Macrobiotus siderophilus		Bertolani et al. 2023
Macrobiotus sottilei	PL.352	MW246133	MW247178	MW247179	MW247175	Macrobiotus sottilei		Kiosya et al. 2021
Macrobiotus sp.	DI02	OP684780		OP696675		Macrobiotus sp.		Amezcua-Martinez et al. (unpubl.)
Macrobiotus sp. 1 MC-2022d	UK 12	OQ525891	OQ525899	OQ548104	OQ525904	Macrobiotus sp. 1 MC-2022d		Mioduchowsk et al. (unpubl.)
Macrobiotus sp. 1 MC-2022d	UK 14	OQ525890	OQ525897		OQ525905	Macrobiotus sp. 1 MC-2022d		Mioduchowsk et al. (unpubl.)
Macrobiotus sp. 1 YW-2020	BN140005	MT863704	MT872214			Macrobiotus sp. 1 YW-2020		Wang (unpubl.)
Macrobiotus sp. 1 ZY-2020	HP1110201	MW187003	MW183923			Macrobiotus sp. 1 ZY-2020		Yuan et al. (unpubl.)
Macrobiotus sp. n. MM-2020	MAD967	MT246661	MT241900		MT241898	Macrobiotus sp. n. MM-2020		Kuzdrowska et al. (unpubl.)
Macrobiotus terminalis	C2868_N02	JN673959	OP596308			Macrobiotus terminalis		Cesari et al. 2011
Macrobiotus vattenrikense sp. nov	22-003	PX093655	PX093663	PX093644	PX093649		COI haplotype 3; 18S haplotype 1; ITS haplotype 1; 28S haplotype 1	this study
Macrobiotus vattenrikense sp. nov	22-004-1	PX093655					COI haplotype 3	this study
Macrobiotus vattenrikense sp. nov	22-017	PX093653		PX093644	PX093649		COI haplotype 1; ITS haplotype 1; 28S haplotype 1	this study
Macrobiotus vattenrikense sp. nov	22-101	PX093654	PX093664	PX093644	PX093650		COI haplotype 2; 18S haplotype 2; ITS haplotype 1; 28S haplotype 2	this study
Macrobiotus vladimiri	FI.068	MN888327	MN888375	MN888347	MN888360	Macrobiotus vladimiri		Vecchi & Stec 2021
Mesobiotus aradasi	KPRI-MF-1	MW727934	MW751962			Mesobiotus cf. furciger	see Vecchi et al. 2024a	Short et al. 2022
Mesobiotus aradasi	S1982.Meb.1	OQ928737	OQ933000	OQ940183	OQ932999	Mesobiotus aradasi		Vecchi et al. 2024a
Mesobiotus bockebodicus	22-044	PQ365775	PQ367888		PQ367894	Mesobiotus sp. 1 SA-2024a	temporary taxon name has not been updated in GenBank at time of writing	Atherton et al. 2025
Mesobiotus cf. barabanovi	KG.002	MN313170	MN310392	MN310390	MN310388	Mesobiotus cf. barabanovi		Kaczmarek et al. 2020a
Mesobiotus cf. furciger	AF01-MF-2	MW727944	MW751946			Mesobiotus cf. furciger		Short et al. 2022
Mesobiotus datanlanicus	VN.026	MK578905	MK584659	MK584657	MK584658	Mesobiotus datanlanicus		Stec 2019
Mesobiotus diegoi	ZA.001.02	OP143857	OP142526	OP142514	OP142520	Mesobiotus diegoi		Stec 2022a
Mesobiotus dilimanensis	PH.006	MN257047	MN257048	MN257050	MN257049	Mesobiotus dilimanensis		Itang et al. 2020
Mesobiotus efa	Lembolovo.DT262	OR803035	OR804457	OR805169	OR805135	Mesobiotus sp. 1 DVT-2023b	temporary taxon name has not been updated in GenBank at time of writing	Tumanov et al. 2024
Mesobiotus emiliae	22-046	PQ365768	PQ367886		PQ367892	Mesobiotus emiliae		Atherton et al. 2025
Mesobiotus ethiopicus	ET.004	MF678794	MF678793	MN122776	MF678792	Mesobiotus ethiopicus		Stec & Kristensen 2017
Mesobiotus fiedleri	MG.001	MH676056	MH681585	MH681724	MH681693	Mesobiotus fiedleri		Kaczmarek et al. 2020a
Mesobiotus furciger	Macro_06_313	MW727960	MW751939			Mesobiotus furciger		Short et al. 2022
Mesobiotus harmsworthi	haplotype 1	MH195150	MH197146	MH197154	MH197264	Mesobiotus harmsworthi		Kaczmarek et al. 2018
Mesobiotus hilariae	Novolazarevskaya_ C_1	KT226108	KT226071			Mesobiotus hilariae		Vecchi et al. 2016
Mesobiotus huecoensis	S2027.4	OQ756247	OQ756248			Mesobiotus huecoensis		Vecchi et al. 2023
Mesobiotus imperialis	VN.061.01	OL311514	OL257855		OL257867	Mesobiotus imperialis		Stec 2021
Mesobiotus insanis		MF 441491	MF441488	MF441490	MF441489	Mesobiotus insanis		Mapalo et al. 2017
Mesobiotus maklowiczi	ZA.002.01	OP143855	OP142525		OP142518	Mesobiotus maklowiczi		Stec 2022a
Mesobiotus mandalori	2.18.1.2	OP825090	OP829143	OP829058	OP829056	Mesobiotus sp. m MM-2022	temporary taxon name has not been updated in GenBank at time of writing	Erdmann et al. 2024
Mesobiotus mandalori	22-053	PQ365769	PQ367887		PQ367893	Mesobiotus sp. m MM-2022	temporary taxon name has not been updated in GenBank at time of writing	Atherton et al. 2025
Mesobiotus marmoreus	VN.055.01	OL311516	OL257856	OL257861	OL257868	Mesobiotus marmoreus		Stec 2022a
Mesobiotus occultatus	DT97	OR803042	OR794157	OR805249	OR794158	Mesobiotus occultatus		Tumanov et al. 2024
Mesobiotus peterseni	GL.002.01	OP143859	OP142528	OP142516	OP142522	Mesobiotus peterseni		Stec 2022a
Mesobiotus philippinicus		KX 129796	KX129793	KX129795	KX129794	Mesobiotus philippinicus		Mapalo et al. 2016
Mesobiotus radiatus	KE.008	MH195147	MH197153	MH197267	MH197152	Mesobiotus radiatus		Stec et al. 2018d
Mesobiotus romani	EC.002	MH195149	MH197158	MH197150	MH197151	Mesobiotus romani		Roszkowska et al. 2018
Mesobiotus skanensis	22-153	PQ365777	PQ367889		PQ367895	Mesobiotus sp. 2 SA-2024a	temporary taxon name has not been updated in GenBank at time of writing	Atherton et al. 2025
Mesobiotus skorackii	CN8	MW656257	MW680643		MW680636	Mesobiotus skorackii		Kayastha et al. 2021
Mesobiotus sp.	BI02	OP684777		OP696672		Mesobiotus sp.		Amezcua-Martinez et al. (unpubl.)
Mesobiotus sp.	NO.234	MH195153	MH197148	MH197156	MH197265	Mesobiotus furciger group sp.		Kaczmarek et al. 2018
Mesobiotus sp.	RU.017	MH195154	MH197149	MH197157	MH197266	Mesobiotus harmsworthi group sp.		Kaczmarek et al. 2018
Mesobiotus sp. 1	DVT-2020	MT904513	MT903468	MT903470	MT903612	Mesobiotus sp. 1		Tumanov 2020
Mesobiotus sp. Fl	Mes_sp_S16_3	OQ968323	OQ974689	OQ974696	OQ974704	Mesobiotus sp. Fl		Vecchi et al. 2024b
Mesobiotus sp. VN	Mes_sp_ VN 036_04	OQ968314	OQ974691			Mesobiotus sp. VN		Vecchi et al. 2024b
Mesobiotus vulpinus	DT553	OR803040	OR804461	OR805172	OR805140	Mesobiotus sp. 2 DVT-2023b	temporary taxon name has not been updated in GenBank at time of writing	Tumanov et al. 2024
Mesobiotus zelmae	22-239	PQ365781	PQ367891		PQ367896	Mesobiotus sp. 3 SA-2024a	temporary taxon name has not been updated in GenBank at time of writing	Atherton et al. 2025
Mesobiotus cf. intermedius	DT270	OP013287	OP035718	OP035707	OP035798	Minibiotus cf. intermedius		Tumanov et al. 2022
Minibiotus citlalium	AI04	OP684765		OP696660		Minibiotus citlalium		Amezcua-Martinez et al. (unpubl.)
Minibiotus furcatus	Tar527	FJ435802	FJ435746		FJ435760	Minibiotus furcatus		Guil & Giribet 2012
Minibiotus gumersindoi	Tar710	FJ435803	FJ435748		FJ435761	Minibiotus gumersindoi		Guil & Giribet 2012
Minibiotus intermedius	Min4 GR	ON005160	ON005189		ON005195	Minibiotus intermedius		Kaczmarek et al. 2022
Minibiotus ioculator	ZA.274	MT023412	MT023998	MT024000	MT024041	Minibiotus ioculator		Stec et al. 2020b
Minibiotus pentannulatus	TZ.027 haplotype 1 MT023413		MT023999	MT024001	MT024042	Minibiotus pentannulatus		Stec et al. 2020b
Minibiotus sidereus	AI12	OP684770		OP696665		Minibiotus sidereus		Amezcua-Martinez et al. (unpubl.)
Minibiotus sp. A MV2020	S69.Min.01	MW306859	OK663227	OK663216	OK663238	Minibiotus sp. A MV2020		Vecchi et al. 2022b
Paramacrobiotus marchelmoni sp. nov.	22-004	PX093656	PX093665	PX093646	PX093651		COI haplotype 1; 18S haplotype 1; ITS haplotype 2; 28S haplotype 1	this study
Paramacrobiotus marchelmoni sp. nov.	22-006	PX093656	PX093665	PX093646			COI haplotype 1; 18S haplotype 1; ITS haplotype 2	this study
Paramacrobiotus marchelmoni sp. nov.	22-007	PX093656		PX093646			COI haplotype 1; ITS haplotype 2	this study
Paramacrobiotus marchelmoni sp. nov.	22-019	PX093656	PX093665	PX093646	PX093651		COI haplotype 1; 18S haplotype 1; ITS haplotype 2; 28S haplotype 1	this study
Paramacrobiotus marchelmoni sp. nov.	22-026	PX093656					COI haplotype 1	this study
Paramacrobiotus marchelmoni sp. nov.	22-029	PX093656		PX093645	PX093651		COI haplotype 1; 18S haplotype 1; 28S haplotype 1	this study
Paramacrobiotus marchelmoni sp. nov.	22-034	PX093656	PX093665	PX093646			COI haplotype 1; 18S haplotype 1; ITS haplotype 2	this study
Paramacrobiotus marchelmoni sp. nov.	22-102	PX093656		PX093645			COI haplotype 1; ITS haplotype 1	this study
Paramacrobiotus marchelmoni sp. nov.	22-106	PX093657					COI haplotype 2	this study
Paramacrobiotus marchelmoni sp. nov.	22-108	PX093656					COI haplotype 1	this study
Paramacrobiotus marchelmoni sp. nov.	22-110	PX093657	PX093665	PX093646	PX093651		COI haplotype 2; 18S haplotype 1; ITS haplotype 2; 28S haplotype 1	this study
Paramacrobiotus marchelmoni sp. nov.	22-113	PX093657	PX093665	PX093646			COI haplotype 2; 18S haplotype 1; ITS haplotype 2	this study
Paramacrobiotus marchelmoni sp. nov.	23-015	PX093658					COI haplotype 3	this study
Paramacrobiotus marchelmoni sp. nov.	23-041	PX093657					COI haplotype 2	this study
Paramacrobiotus aff. arduus	PRODO-3	MK041022				Paramacrobiotus aff. arduus		Guidetti et al. 2019a
Paramacrobiotus aff. spatialis OLBIA-1		MK041002	MK041026			Paramacrobiotus aff. spatialis		Guidetti et al. 2019a
Paramacrobiotus arduus	PRODO-1	MK041020	MK041032			Paramacrobiotus arduus		Guidetti et al. 2019a
Paramacrobiotus arduus	PRODO-2	MK041021				Paramacrobiotus arduus		Guidetti et al. 2019a
Paramacrobiotus areolatus	NO.385	MH675998	MH664931	MH666080	MH664948	Paramacrobiotus areolatus		Stec et al. 2020c
Paramacrobiotus bifrons	C4497_ US 1	PP236542		PP240910		Paramacrobiotus sp. 1 MC-2024a	temporary taxon name has not been updated in GenBank at time of writing	Brandoli et al. 2024
Paramacrobiotus bifrons	C4497_ US 2	PP236543		PP240911		Paramacrobiotus sp. 1 MC-2024a	temporary taxon name has not been updated in GenBank at time of writing	Brandoli et al. 2024
Paramacrobiotus bifrons	C4497_ US 7	PP236544		PP240912		Paramacrobiotus sp. 1 MC-2024a	temporary taxon name has not been updated in GenBank at time of writing	Brandoli et al. 2024
Paramacrobiotus bifrons	C4497_ US 8	PP236545		PP240913		Paramacrobiotus sp. 1 MC-2024a	temporary taxon name has not been updated in GenBank at time of writing	Brandoli et al. 2024
Paramacrobiotus bifrons	C5025_ US 6	PP236551		PP240918		Paramacrobiotus sp. 1 MC-2024a	temporary taxon name has not been updated in GenBank at time of writing	Brandoli et al. 2024
Paramacrobiotus bifrons	C5062_ US 2	PP236553		PP240919		Paramacrobiotus sp. 1 MC-2024a	temporary taxon name has not been updated in GenBank at time of writing	Brandoli et al. 2024
Paramacrobiotus bifrons	C5062_ US 3	PP236554		PP240920		Paramacrobiotus sp. 1 MC-2024a	temporary taxon name has not been updated in GenBank at time of writing	Brandoli et al. 2024
Paramacrobiotus bifrons	IT.048	MH676007	MH664937	MH666085	MH664955	Paramacrobiotus sp.	see Brandoli et al. 2024	Stec et al. 2020c
Paramacrobiotus bifrons	Par_cf_kly_ S108_2	OQ968322		OQ974697		Paramacrobiotus cf. klymenki	see Brandoli et al. 2024	Vecchi et al. 2024b
Paramacrobiotus celsus	PIANE_DI_ MOCOGNO-1	MK041017	MK041031			Paramacrobiotus celsus		Guidetti et al. 2019a
Paramacrobiotus celsus	PIANE_DI_ MOCOGNO-2	MK041018				Paramacrobiotus celsus		Guidetti et al. 2019a
Paramacrobiotus celsus	PIANE_DI_ MOCOGNO-3	MK041019				Paramacrobiotus celsus		Guidetti et al. 2019a
Paramacrobiotus danielae	GY26001		MZ081363	MZ081364		Paramacrobiotus danielae		Wang et al. (unpubl.)
Paramacrobiotus depressus	ANDALO-1	MK041015				Paramacrobiotus depressus		Guidetti et al. 2019a
Paramacrobiotus depressus	OSPITALETTO_ A-1	MK041016				Paramacrobiotus depressus		Guidetti et al. 2019a
Paramacrobiotus depressus	PASSO_ BALLINO-1	MK041012	MK041030			Paramacrobiotus depressus		Guidetti et al. 2019a
Paramacrobiotus depressus	PASSO_ BALLINO-2	MK041013				Paramacrobiotus depressus		Guidetti et al. 2019a
Paramacrobiotus depressus	PASSO_ BALLINO-3	MK041014				Paramacrobiotus depressus		Guidetti et al. 2019a
Paramacrobiotus experimentalis	MAD-TAR11	MN097837		MN073464	MN073465	Paramacrobiotus experimentalis		Kaczmarek et al. 2020c
Paramacrobiotus fairbanksi	22-423	PX093660	PX093666			Paramacrobiotus fairbanksi		this study
Paramacrobiotus fairbanksi	23-022	PX093661				Paramacrobiotus fairbanksi		this study
Paramacrobiotus fairbanksi	23-041	PX093659	PX093666	PX093647	PX093648	Paramacrobiotus fairbanksi		this study
Paramacrobiotus fairbanksi	Alb1	ON911918				Paramacrobiotus fairbanksi		Mioduchowsk et al. (unpubl.)
Paramacrobiotus fairbanksi	Alb4	ON911917				Paramacrobiotus fairbanksi		Mioduchowsk et al. (unpubl.)
Paramacrobiotus fairbanksi	C2345-MR_2	AY598778				Macrobiotus richtersi		Guidetti et al. 2005
Paramacrobiotus fairbanksi	C2345-MR_3	AY598779				Macrobiotus richtersi		Guidetti et al. 2005
Paramacrobiotus sp.	Kenya 3	EU244599				Paramacrobiotus sp. ‘ richtersi ’ group		Schill et al. 2010
Paramacrobiotus sp.	MG.002	MH676008	MH664938	MH666086	MH664956	Paramacrobiotus sp.		Stec et al. 2020c
Paramacrobiotus sp.	NO.386	MH676009	MH664939	MH666088	MH664957	Paramacrobiotus sp.		Stec et al. 2020c
Paramacrobiotus sp.	NZ.001	MH676010	MH664940	MH666089	MH664958	Paramacrobiotus sp.		Stec et al. 2020c
Paramacrobiotus sp.	Par_sp_S175_2	OQ968321				Paramacrobiotus sp.		Vecchi et al. 2024b
Paramacrobiotus sp.	PT.006	MH676013	MH664943	MH666092	MH664960	Paramacrobiotus sp.		Stec et al. 2020c
Paramacrobiotus sp.	PT.048 haplotype 1	MH676014	MH664944	MH666093	MH664961	Paramacrobiotus sp.		Stec et al. 2020c
Paramacrobiotus sp.	PT.048 haplotype 2	MH676015				Paramacrobiotus sp.		Stec et al. 2020c
Paramacrobiotus sp.	T3	OP531839	ON923868			Paramacrobiotus sp.		Basu et al. 2023
Paramacrobiotus sp.	Tar407	FJ435807	FJ435741		FJ435756	Macrobiotus pallarii	see Discussion	Guil & Giribet 2012
Paramacrobiotus sp.	TN.014	MH676016	MH664945	MH666094	MH664962	Paramacrobiotus sp.		Stec et al. 2020c
Paramacrobiotus sp.	TZ.018	MH676017	MH664933	MH666095	MH664951	Paramacrobiotus sp.		Stec et al. 2020c
Paramacrobiotus sp. DLS-2024a	T82_S135	PP414782	PP416751	PP416753	PP416752	Paramacrobiotus sp. DLS-2024a		López-Sandoval et al. 2025
Paramacrobiotus sp. MAC-2014	T1	KF788251				Paramacrobiotus richtersi group sp. 1 MAC-2014		Caicedo et al. 2017
Paramacrobiotus sp. MAC-2014	T2	KF788252				Paramacrobiotus richtersi group sp. 1 MAC-2014		Caicedo et al. 2017
Paramacrobiotus sp. MAC-2014	T3	KF788253				Paramacrobiotus richtersi group sp. 1 MAC-2014		Caicedo et al. 2017
Paramacrobiotus sp. MAC-2014	T4	KF788254				Paramacrobiotus richtersi group sp. 1 MAC-2014		Caicedo et al. 2017
Paramacrobiotus sp. MAC-2014	T6	KF788255				Paramacrobiotus richtersi group sp. 1 MAC-2014		Caicedo et al. 2017
Paramacrobiotus sp. MAC-2014	T7	KF788256				Paramacrobiotus richtersi group sp. 1 MAC-2014		Caicedo et al. 2017
Paramacrobiotus sp. MAC-2014	T8	KF788257				Paramacrobiotus richtersi group sp. 1 MAC-2014		Caicedo et al. 2017
Paramacrobiotus sp. ZY-2020	hp1100404	MT731035	MT723894			Paramacrobiotus sp. ZY-2020		Yuan et al. (unpubl.)
Paramacrobiotus sp. ZY-2022c	N 3	OP870172			ON819001	Paramacrobiotus sp. ZY-2022c		Yuan et al. (unpubl.)
Paramacrobiotus spatialis	FORMIGINE-1	MK040995	MK041024			Paramacrobiotus spatialis		Guidetti et al. 2009
Paramacrobiotus spatialis	FORMIGINE-2	MK040996				Paramacrobiotus spatialis		Guidetti et al. 2019a
Paramacrobiotus spatialis	FORMIGINE-3	MK040997				Paramacrobiotus spatialis		Guidetti et al. 2019a
Paramacrobiotus spatialis	FORMIGINE-4	MK040998				Paramacrobiotus spatialis		Guidetti et al. 2019a
Paramacrobiotus spatialis	GAGGIO-1	MK040999				Paramacrobiotus spatialis		Guidetti et al. 2019a
Paramacrobiotus spatialis	GAGGIO-2	MK041000				Paramacrobiotus spatialis		Guidetti et al. 2019a
Paramacrobiotus spatialis	GAGGIO-3	MK041001	MK041025			Paramacrobiotus spatialis		Guidetti et al. 2019a
Paramacrobiotus spatialis	S107.Par	OK662996	OK663225	OK663214	OK663236	Paramacrobiotus spatialis		Vecchi et al. 2022c
Paramacrobiotus tonollii	US.035	MH676018	MH664946	MH666096	MH664963	Paramacrobiotus tonollii		Stec et al. 2020c
Richtersius coronifer	NO.385	MH676053	MH681760	MH681763	MH681757	Richtersius coronifer		Stec et al. 2020c
Sisubiotus hakaiensis	Sis.hak.S1911	OM523181	OM523054	OM523057	OM523059	Sisubiotus hakaiensis		Vecchi et al. 2022a
Sisubiotus spectabilis	FI.067	MN888322	MN888371	MN888331	MN888357	Sisubiotus spectabilis		Stec et al. 2021a
Tenuibiotus cf. ciprianoi	ES.086	MN888328	MN888376	MN888348	MN888361	Tenuibiotus cf. ciprianoi		Stec et al. 2021a
Tenuibiotus danilovi	KG.128	MN888329	MN888377	MN888349	MN888362	Tenuibiotus danilovi		Stec et al. 2021a
Tenuibiotus sp.	22-109	PX093662	PX093667		PX093652	Tenuibiotus sp.		this study
Tenuibiotus tenuiformis	KG.140	MN888330	MN888378	MN888350	MN888363	Tenuibiotus tenuiformis		Stec et al. 2021a
Tenuibiotus voronkovi	haplotype 1	KX810042	KX810045	KX810046	KX810049	Tenuibiotus voronkovi		Zawierucha et al. 2016
Tenuibiotus yeliseii	DT422	OR145334	OR142418	OR142424	OR142426	Tenuibiotus sp. 1 DVT-2023a	temporary taxon name has not been updated in GenBank at time of writing	Tsvetkova & Tumanov 2024
Tenuibiotus zandrae		MN 444827	MN443040	MN443038	MN443035	Tenuibiotus zandrae		Stec et al. 2020c
Xerobiotus arenosum	C4411_V4	OR397039	OR398024		OR398050	Xerobiotus arenosum		Vincenzi et al. 2024
Xerobiotus cf. reductus	PL3177_1	OR397050	OR398033		OR398056	Xerobiotus cf. reductus		Vincenzi et al. 2024
Xerobiotus euxinus	check 1.4	PP469626	PP439986	PP470593	PP440001	Macrobiotus euxinus	see Vincenzi et al. 2024	Polishchuk et al. 2024
Xerobiotus euxinus	C3834_V4	OR397005	OR398005		OR398119	Macrobiotus euxinus	see Vincenzi et al. 2024	Vincenzi et al. 2024
Xerobiotus gretae	C4341 G	MW581665	MW588434	MW588431	MW588438	Macrobiotus gretae	see Vincenzi et al. 2024	Massa et al. 2021
Xerobiotus litus	C4383_2	OR397000	OR398002		OR398118	Xerobiotus litus		Vincenzi et al. 2024
Xerobiotus naginae	S226.Mac.1	OK662990	OK663219		OK663230	Marobiotus naginae	see Vincenzi et al. 2024	Vecchi et al. 2022c
Xerobiotus pseudohufelandi	C2358-XP1	AY598776	HQ604989			Xerobiotus pseudohufelandi		Guidetti et al. 2005
Xerobiotus reductus	PL_3176_1	OR397048	OR398028		OR398130	Xerobiotus reductus		Vincenzi et al. 2024
Xerobiotus sp.	PL.360	MN888325	MN888373	MN888345	MN888358	Xerobiotus sp.		Vecchi & Stec 2021
Xerobiotus sp.	ZA.373	MN888326	MN888374	MN888346	MN888359	Xerobiotus sp.		Vecchi & Stec 2021
Xerobiotus sp. 1 MC-2023a	C3306_V1	OR397025	OR398020		OR398123	Xerobiotus sp. 1 MC-2023a		Vincenzi et al. 2024