| Body length | 119.1–129.7 | 89.0–140.0 | 141.0–173.0 |
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| Cephalic pleuria | Cephalion narrow, extended dorsally; epipleuria small visible from the dorsal body side; hypopleuria large not visible from dorsal body side. Hypostomium present | Cephalion wide; epipleuria large; hypopleuria small; clearly visible from the dorsal body side. Hypostomium present | Cephalion wide; epipleuria large, clearly visible on the dorsal body side; hypopleuria smallpresent only on the ventral body side. Hypostomium absent |
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| Place of arising cephalic cilia | Anterior tufts: in lines between cephalion and epipleuria; posterior tufts: at the anterior edge of the hypopleuria on the ventral side | Anterior and posterior tufts: lateral | Anterior tufts: dorsally at the head top; posterior: dorsally |
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| Pharynx length | 31.9–34.9 | 26.0–33.0 | 36.0–44.0 |
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| Anterior intestinal section morphologically different | Present | Absent | Absent |
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| Furca length | 19.8–23.2 | 12.0–17.0 | 14.0–20.0 |
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| Adhesive tube length | 10.1–11.3 | 9.0 | 7.0–11.0 |
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| Type of furca indentation | V-shaped | V-shaped | Helmet-shaped |
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| Number of scales in single longitudinal row | 24–27 | 20 | 25–33 |
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| Total number of longitudinal alternating rows of scales | 29–35 | 23–31 | 45–53 |
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| Type of scales | Almost all three-lobed with strong keels and straight spines; shape strongly differentiated within body regions | One-lobed with strong keels and straight spines; shape undifferentiated within body regions | One-lobed with strong keels and straight spines; shape undifferentiated within body regions |
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| Size of scales | Small; size strongly differentiated within body regions | Small; size slightly increasing posteriorly | Small; similar size all over the body |
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| Presence of scales with two keels and two spines on the furcal appendages | Absent | One pair of scales with two keels and two spines present dorsolateraly | Absent |
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| Scale coverage of the ventral interciliary field | Only the posterior end of the trunk covered by scales | Only the posterior end of the trunk covered by scales | Entire ventral interciliary field covered by scales |
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| Type of terminal scales of the ventral interciliary field | Oval with shallow posterior notches, keeled, spineless | Oval without posterior notches, keeled, spined | Oval without posterior notches, keeled, spineless |
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| Variety of spines length | Strongly differentiated length of spines on the head, neck (short, vestigial on dorsal area) and trunk; length of the spines gradually lengthen from the lateral to the ventral sides of the body; much longer spines on the ventral body sides | Similar length of spines on the dorsal, dorsolateral, lateral sides of the head, neck and trunk; much longer spines on the ventrolateral body sides | All spines similar length |
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| Type of posteriormost pair of the lateral trunk spines | Slightly longer and thicker than surrounding spines | Same as other spines | Same as other spines |
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| Type of last pair of the parafurcal spines | Longer, thicker and stronger than other spines | Same as other spines | Same as other spines |
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| Number of dorsal sensory bristles | 3 pairs | 2 pairs | 2 pairs |
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