Systematics of the Calotes irawadi complex (Squamata, Agamidae) with two newly described species from Thailand Author Prakobkarn, Arpapan https://orcid.org/0009-0000-9047-1102 Ph. D. Program in Zoology, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Zug, George R. 0000-0003-4922-5721 Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, USA Author Tandavanitj, Nontivich Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Ngamprasertwong, Thongchai 0000-0003-3146-1849 Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand text ZooKeys 2026 2026-06-03 1281 69 104 journal article 315339 10.3897/zookeys.1281.175455 1321a16e-cff9-4247-8200-6557d35c47d3 E54EA084-1419-444B-B507-C4646540A63B

Table 6. Spine characters of male Calotes thailandensis sp. nov. and male C. maehongsonensis sp. nov. Measurements are represented in mean ± SD (range) in millimeters (mm). Abbreviations are showed in Materials and methods.

Characters	Calotes thailandensis sp. nov.		Calotes maehongsonensis sp. nov.
	Holotype	Paratypes	Holotype	Paratypes
	CUMZ-R -2738	Males (n = 5)	CUMZ-R -2825	Males (n = 5)
MCS	4.5	4.41 ± 0.90	3.45	4.04 ± 0.63
		(3.60–5.60)		(3.25–4.80)
ASS	2.1	2.30 ± 0.36	1.35	1.10 ± 0.11
		(2.10–2.95)		(1.00–1.25)
PSS	1.85	2.22 ± 0.17	1.2	1.14 ± 0.13
		(2.05–2.40)		(1.00–1.25)
MCS / TymD	1.34	1.33 ± 0.28	1.27	1.44 ± 0.16
		(1.03–1.68)		(1.20–1.62)
ASS / TymD	0.61	0.69 ± 0.08	0.5	0.39 ± 0.02
		(0.59–0.82)		(0.37–0.42)
PSS / TymD	0.55	0.67 ± 0.03	0.44	0.41 ± 0.06
		(0.64–0.70)		(0.33–0.47)