A new species in the Hipposideros bicolor group (Chiroptera: Hipposideridae) from Peninsular Malaysia Author Murray, Susan W. Center for Ecology and Conservation Biology Department of Biology Boston University Boston MA 02215 USA & Corresponding author: E-mail: susan. m 88 @ gmail. com susan.m88@gmail.com Author Khan, Faisal A. A. Faculty of Resource Science and Technology Universiti Malaysia Sarawak 94300 Kota Samarahan Sarawak Malaysia Author Kingston, Tigga Department of Biological Sciences Texas Tech University Lubbock TX 79409 USA Author Zubaid, Akbar Faculty of Science and Technology Universiti Kebangsaan Malaysia Bangi Selangor Malaysia Author Campbell, Polly Department of Integrative Biology Oklahoma State University Stillwater OK 74078 USA text Acta Chiropterologica 2018 20 1 1 29 journal article 21482 10.3161/15081109ACC2018.20.1.001 e95dc0f3-4897-4a17-aaf4-ce56c28f49f0 1733-5329 3944846

TABLE 2. Mean values for external and wing morphology with standard deviation and sample size (in parentheses) for H. bicolor (= H. bicolor -131) and H. kunzi (H. bicolor -142) captured in peninsular Malaysia (not including bats from Taiping and Kuala Atok). Ranges are also presented for external morphology and echolocation. Values for H. cineraceus -B are included for comparison because of this lineage’s close phylogenetic relationship with H. kunzi, but were not included in statistical analyses. The results of univariate GLM analyses for an effect of species on external measurements and echolocation call frequency with phonic type split by sex are given: * — P <0.05, ** — P <0.01, and *** — P < 0.001. For the wing morphology data, the results of a GLM analysis with the sexes pooled, species as a fixed factor and length of forearm as a covariate are also shown. Differences between males and females within each phonic type were evaluated using t -tests; α = 0.004 after Bonferroni correction

Variable	H. bicolor KWR	H. kunzi	H. cineraceus -B
† — Data from Kingston et al. (2000)
Forearm length (mm)***	45.3 ± 1.0 (143)	42.9 ± 0.9 (864)	39.3 (1)
Range	43.0–48.2	38.8–45.6
♀♀	45.8 ± 0.8 (76)***	43.3 ± 0.8 (414)***
♂♂	44.7 ± 0.8 (67)	42.6 ± 0.8 (450)
Tibia length (mm)***	20.5 ± 0.6 (131)	18.8 ± 0.5 (742)	17.2 (1)
Range	18.7–22.1	17.1–20.4
♀♀	20.7 ± 0.6 (67)***	18.8 ± 0.5 (363)
♂♂	20.3 ± 0.5 (64)	18.8 ± 0.5 (379)
Tail length (mm)***	31.2 ± 2.1 (131)	28.7 ± 1.8 (719)	34.0 (1)
Range	26.0–35.0	21.0–34.0
♀♀	32.0 ± 1.8 (68)***	29.3 ± 1.6 (356)***
♂♂	30.2 ± 2.1 (63)	28.2 ± 1.7 (363)
Ear height (mm)***	18.2 ± 0.7 (132)	17.6 ± 0.6 (666)	16.0 (1)
Range	15.1–20.5	15.0–19.5
♀♀	18.2 ± 0.8 (66)	17.7 ± 0.6 (315)***
♂♂	18.1 ± 0.7 (66)	17.5 ± 0.6 (351)
Body mass (g)**	8.6 ± 0.8 (124)	8.5 ± 0.9 (805)	6.0 (1)
Range	6.8–10.8	5.8–12.0
♀♀	8.9 ± 0.8 (62)***	8.5 ± 1.0 (357)
♂♂	8.3 ± 0.7 (62)	8.5 ± 0.9 (448)
Call frequency (kHz)***	131.8 ± 2.0 (136)	143.1 ± 2.0 (842)	144.0†
Range	124.9–135.8	135.8–147.5
♀♀	132.4 ± 1.4 (75)***	142.8 ± 2.0 (403)***
♂♂	131.0 ± 2.2 (61)	143.4 ± 1.9 (439)
Wing area (m2)	0.0135 ± 0.0008	0.0121 ± 0.0007
♀♀	0.0139 ± 0.0007*	0.0123 ± 0.0007
♂♂	0.0132 ± 0.0007	0.0119 ± 0.0007
Arm-Wing area (m2)**	0.0033 ± 0.0002	0.0031 ± 0.0002
♀♀	0.0034 ± 0.0003	0.0032 ± 0.0002***
♂♂	0.0033 ± 0.0002	0.0030 ± 0.0002
Hand-Wing area (m2)***	0.0024 ± 0.0001	0.0021 ± 0.0001
♀♀	0.0025 ± 0.0001	0.0021 ± 0.0001
♂♂	0.0024 ± 0.0001	0.0021 ± 0.0001
Wing span (m)	0.2863 ± 0.0088	0.2736 ± 0.0088
♀♀	0.2896 ± 0.0079	0.2758 ± 0.0076
♂♂	0.2837 ± 0.0088	0.2715 ± 0.0092
Arm-Wing length (m)**	0.0572 ± 0.0020	0.0549 ± 0.0020
♀♀	0.0577 ± 0.0018	0.0556 ± 0.0018**
♂♂	0.0567 ± 0.0021	0.0542 ± 0.0019
Hand-Wing length (m)*	0.0654 ± 0.0026	0.0629 ± 0.0024
♀♀	0.0664 ± 0.0019	0.0633 ± 0.0023
♂♂	0.0646 ± 0.0029	0.0624 ± 0.0023
Wing loading (N/m2)***	6.19 ± 0.56	6.74 ± 0.76
♀♀	6.32 ± 0.57	6.65 ± 0.76**
♂♂	6.10 ± 0.54	6.84 ± 0.75
Aspect ratio*	6.08 ± 0.23	6.18 ± 0.23
♀♀	6.04 ± 0.21	6.17 ± 0.22
♂♂	6.11 ± 0.24	6.19 ± 0.24
Tip index***	1.76 ± 0.30	1.44 ± 0.28
♀♀	1.72 ± 0.32	1.39 ± 0.24**
♂♂	1.80 ± 0.29	1.49 ± 0.32