Taxonomic review of the genus Johnius (Perciformes: Sciaenidae) from East Malaysian waters, with comments on newly recorded Johnius species Author Hanafi, Norhafiz Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan, R. O. C. Author Chen, Meng-Hsien Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan, R. O. C. & Institute of Marine Ecology and Conservation, National Sun Yat-sen University, Kaohsiung, Taiwan, R. O. C. Author Chang, Chih-Wei Department of Oceanography (Marine Biology group), National Sun Yat-sen University, Kaohsiung, Taiwan, R. O. C. & Marine Ecology and Conservation Research Center, National Academy of Marine Research, Kaohsiung, Taiwan, R. O. C. & National Museum of Marine Biology and Aquarium, and Graduate Institute of Marine Biology, National Dong Hwa University, Pingtung, Taiwan, R. O. C. Author Chao, Ning Labbish National Museum of Marine Biology and Aquarium, and Graduate Institute of Marine Biology, National Dong Hwa University, Pingtung, Taiwan, R. O. C. & Bio-Amazonia Conservation International, Boston, Massachusetts, USA Author Seah, Ying Giat Faculty of Fisheries and Food Science, University Malaysia Terengganu (UMT), Kuala Terengganu, Malaysia & Fish Division, South China Sea Repository and Reference Center, Institute of Oceanography and Environment, University Malaysia Terengganu (UMT), Kuala Terengganu, Malaysia text Zootaxa 2024 2024-12-10 5550 1 85 117 https://doi.org/10.11646/zootaxa.5550.1.11 journal article 306734 10.11646/zootaxa.5550.1.11 0522af6a-1e4c-4918-8940-a921577dee94 1175-5326 14388193 44C6949D-67DE-4C01-ABC8-3FB0F163BF5F

TABLE 6. Morphometric measurement of J. amblycephalus, J. belangerii, J. borneensis, J. carouna and J. coitor.

SPECIES	J. amblycephalus	J. belangerii	J. borneensis	J. carouna	J. coitor
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	(n=31)	(n=17)	(n=30)	(n=30)	(n=4)
Morphometric measurements
Standard length(mm)	106–156	82–157	86–185	88–148	108–136
As percentage of SL
Head length	28.4–32.1(30.4±1.0)	28.6–32.5(30.4±1.3)	29.5–34.3(32.0±1.1)	25.5–32.3(29.83±1.4)	28.9–30.2(2935±0.6)
Pectoral fin length	9.2–34.9(21.8±3.7)	17.3–19.6(18.6±0.6)	12.8–24.1(19.0±2.2)	13.12–26.2(21.5±2.6)	20.7–22.7(21.5±0.8)
Pelvic fin length	14.7–32.2(17.4±2.9)	16.6–21.0(18.8±1.3)	12.2–19.0(16.5±1.8)	13.5–19.9(16.8±1.5)	18.9–21.8(20.1±1.3)
Body depth	22.6–28.1(25.9±1.4)	27.3–32.2(29.2±1.5)	25.8–32.3(29.2±1.5)	20.0–31.6(26.5±2.1)	26.3–27.6(27.1±0.6)
Body width	13.9–19.2(16.1±1.3)	13.0–16.0(14.7±0.8)	11.3–19.1(15.5±1.5)	10.4–19.4(14.1±1.6)	14.4–17.1(15.5±1.2)
1 st dorsal fin base length	18.5–25.6(22.0±1.9)	18.2–23.5(20.6±1.5)	15.2–25.1(20.7±1.8)	15.9–26.9(20.3±2.0)	19.5–22.6(20.4±1.5)
4 th spine of 1 st dorsal fin length	16.2–27.9(21.3±2.7)	11.3–17.7(15.6±1.5)	10.5–17.9(14.8±1.6)	11.3–18.8(15.2±1.7)	14.3–17.5(15.9±1.4)
2 nd dorsal fin base length	31.1–40.4(34.7±2.7)	37.5–46.5(41.2±2.2)	32.1–42.4(38.2±2.4)	35.1–43.7(40.1±2.1)	38.3–41.9(40.5±1.8)
5 th ray of 2 nd dorsal fin length	6.2–11.8(9.2±1.4)	7.8–12.2(10.1±1.2)	6.8–14.7(10.1±1.7)	7.9–17.3(10.4±1.5)	9.1–11.2(10.3±1.0)
Anal fin base length	8.4–11.0(9.4±0.7)	8.7–12.0(10.2±1.0)	7.0–10.4(8.8±0.7)	7.4–10.8(8.8±0.7)	9.2–10.6(9.9±0.6)
2 nd spine of anal fin length	6.3–10.8(8.5±1.1)	8.9–13.4(11.3±1.2)	5.6–13.6(8.4±1.6)	9.6–14.7(11.9±1.3)	10.3–11.6(11.1±0.6)
1 st ray of anal fin length	7.9–14.0(12.4±1.3)	13.0–16.0(14.4±0.7)	8.0–16.6(12.4±1.8)	9.7–16.5(13.2±1.3)	15.2–15.4(15.3±0.1)
Preanal length	61.8–69.8(66.9±1.7)	65.8–69.3(67.8±1.2)	64.2–78.6(71.6±2.3)	62.5–72.7(68.3±2.0)	63.3–67.0(65.8±1.7)
Predorsal length	32.3–37.0(34.7±1.2)	33.3–37.1(34.7±1.2)	32.8–36.7(35.2±1.0)	27.0–38.0(34.0±2.1)	34.5–36.5(35.8±0.9)
Snout to origin of 2 nd dorsal fin length	52.2–59.0(54.6±1.7)	50.3–57.3(53.1±1.8)	51.7–57.9(54.4±1.5)	43.6–57.5(52.3±2.5)	52.0–56.7(54.5±2.1)
Prepectoral length	28.1–38.1(31.4±1.8)	28.0–31.4(29.6±0.9)	29.2–34.6(32.0±1.3)	22.7–34.3(30.1±2.2)	30.9–32.0(31.3±0.5)
Prepelvic length	27.8–36.3(33.5±1.7)	32.7–35.1(33.9±0.6)	33.2–38.1(36.0±1.3)	27.0–34.3(30.1±2.2)	33.5–35.5(34.4±0.8)
Caudal peduncle depth	7.6–10.1(9.0±0.7)	7.6–10.1(9.7±0.6)	7.8–10.9(9.8±0.8)	2.2–10.5(9.2±1.1)	7.6–7.9(7.8±0.2)
As percentage of HL
Snout length	25.9–34.0(30.3±2.1)	24.5–29.2(27.1±1.4)	22.3–33.4(29.6±2.1)	23.8–30.2(27.1±1.5)	32.1–33.6(32.6±0.7)
Upper jaw length	35.2–56.1(48.7±5.4)	35.2–50.5(42.1±4.3)	36.9–56.0(46.7±5.5)	34.7–52.6(43.6±5.7)	42.3–45.6(43.8±1.4)
Tip upper jaw to hinge length	26.1–38.0(32.9±3.5)	24.8–37.1(29.6±4.0)	28.2–46.9(37.5±4.8)	22.6–39.8(30.4±5.0)	27.1–29.9(28.9±1.3)
Tip lower jaw to hinge length	19.5–33.6(28.1±3.5)	22.0–33.3(26.5±3.5)	19.8–42.7(33.8±5.1)	21.0–46.4(27.2±5.2)	24.8–27.5(26.0±1.1)
Eye diameter	19.2–27.1(22.8±1.9)	20.8–32.6(25.3±3.4)	19.3–27.3(22.6±1.6)	20.3–35.2(24.8±2.5)	12.3–16.5(14.1±1.8)
Interorbital width	26.7–36.1(29.1±1.6)	18.5–24.4(20.8±2.1)	24.9–31.8(27.3±1.6)	20.9–29.2(25.1±1.8)	26.2–28.0(27.0±0.9)