Acoustic signals, species diagnosis, and species concepts: the case of a new cryptic species of Leptodactylus (Amphibia, Anura, Leptodactylidae) from the Chapare region, Bolivia Author Angulo, Ariadne Author Reichle, Steffen text Zoological Journal of the Linnean Society 2008 2008-03-31 152 3 59 77 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00447.x journal article 3192 10.1111/j.1096-3642.2008.00447.x a85e8f06-95fa-467d-8d09-c49f593271a3 0024-4082 4687663

Table 3. Morphometric measurements for adult specimens of described species of the Leptodactylus marmoratus group. Single numbers refer to holotype measurements when only these are available (except for the case of Leptodactylus diptyx, where five undesignated specimens were listed in the species description, but for which no ranges were provided); numbers in brackets refer to ranges derived from type series measurements, except for Leptodactylus nanus (measurements provided in Kwet, 2007). All measuments are expressed in mm. Measurements for Leptodactylus hylaedactylus were converted from lines (units of measurement equivalent to one-twelfth of an inch)

Species	SVL	HL	HW	ED	TYD	END	IOD	IND	THL	FL	Source
*While there are several specimens in the type species, the author provides measurements only for the holotype and the largest female. †Although there are other type specimens for Leptodactylus bokermanni, measurements for these were not included as they come from different localities and could comprise different species. ‡A more recent study provides new data on morphological variation (Kok et al., 2007).
Leptodactylus andreae *	(20–21)	(8–8.5)	(7.5–9)	NA	½ of ED	NA	NA	NA	NA	NA	Müller (1923)
Leptodactylus araucaria (n = 11)	(17.1–19.9)	(5.7–6.9)	(5.5–6.7)	(1.7–2.0)	(1.1–1.3)	(1.4–1.7)	3.4–3.9)	(1.8–2.2)	(7.3–8.2)	(9.0–10.1)	Kwet & Angulo (2002)
Leptodactylus bokermanni †	24.0	9.0	7.9	2.7	1.3	2.0	2.2	NA	9.7	12.3	Heyer (1973)
Leptodactylus diptyx (n = 5)	22	NA	NA	NA	½ of ED	NA	NA	NA	NA	NA	Boettger (1885)
Leptodactylus heyeri (n = 4)	(22.5–25.8)	(9.1–11.0)	(9.2–10.7)	(2.3–3.0)	(1.5–2.0)	(1.9–2.0)	(2.5–3.3)	(2.1–2.7)	NA	NA	Boistel et al. (2006)
Leptodactylus hylaedactylus (n = 1)	24.6	NA	8.5	NA	NA	NA	NA	NA	NA	NA	Cope (1868)
Leptodactylus lutzi ‡ (n = 3)	(30.1–34.1)	11.1	11.2	3.0	1.7	2.8	2.4	NA	13.4	14.9	Heyer (1975)
Leptodactylus marmoratus (n = 1)	21.6	7.7	7.4	NA	NA	NA	NA	NA	9.2	NA	Kwet & Angulo (2002)
Leptodactylus martinezi (n = 2)	(23–23)	NA	(7–7)	(2.6–2.7)	(1.0–1.0)	(2.2–2.3)	NA	(2.0–2.1)	NA	NA	Bokermann (1956)
Leptodactylus nanus (n = 25)	(16.3–20)	(5.5–6.8)	(5.4–6.4)	(1.4–2.0)	(0.9–1.4)	(1.4–2.1)	(3.2–3.9)	(1.7–2.3)	(7.4–9.6)	(8.5–10.4)	Kwet (2007)
Leptodactylus thomei (n = 25)	(19.18–23.21)	(5.19–6.38)	(5.50–6.75)	(1.92–2.40)	(0.92–1.20)	(1.52–1.88)	(1.69–2.06)	(1.88–2.36)	(6.25–8.63)	(7.13–9.50)	Almeida & Angulo (2006)