Acoustic signals, species diagnosis, and species concepts: the case of a new cryptic species of Leptodactylus (Amphibia, Anura, Leptodactylidae) from the Chapare region, Bolivia Author Angulo, Ariadne Author Reichle, Steffen text Zoological Journal of the Linnean Society 2008 2008-03-31 152 3 59 77 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00447.x journal article 3192 10.1111/j.1096-3642.2008.00447.x a85e8f06-95fa-467d-8d09-c49f593271a3 0024-4082 4687663

Table 2. Measurement of advertisement call parameters for Leptodactylus coca sp. nov. Numbers in the first line are means ± standard deviation; numbers in brackets are ranges

Recording	*NKA 3630	CBF 2619
* Holotype.
Number of calls	23	18
Temperature (°C)	20.8	21.1
Call length (ms)	117.96 ± 3.2 (110.14–123.45)	141.87 ± 2.51 (136.03–145.22)
Call rate (calls/min)	84	66
Call rise time (ms)	52.08 ± 22.17 (23.63–88.62)	41.71 ± 20.25 (11.56–96.05)
Pulses/call	12 ± 1.14 (10–15)	Strong amplitude modulation,
		but no distinct pulses
Pulse rate (/s)	120 ± 12.85 (105.78–146.88)	Strong amplitude modulation,
		but no distinct pulses
Fundamental frequency (Hz)	1731.9 ± 33.11 (1694.6–1794.3)	1855.2 ± 26 (1814.2–1913.9)
Dominant frequency (Hz)	3599.03 ± 68.5 (3449.1–3668.4)	3678.3 ± 29.2 (3628.5–3748.1)
Change in dominant frequency (Hz)	254.65 ± 173.2 (0–775.2)	445.02 ± 99.10 (258.4–602.99)
Total number of harmonics detected	Up to 4	Up to 5 or 6