Five new earthworm species of the Pheretima sangirensis-group (Clitellata: Megascolecidae) from Mount Banahaw, Quezon Province, Philippines Author Hong, Yong 0000-0002-8093-9717 Department of Plant Medicine, College of Agriculture & Life Sciences, Jeonbuk National University, Jeonju 54896, Korea yonghong@jbnu.ac.kr Author James, Samuel W. 0000-0001-6758-5726 Department of Regenerative Organic Agriculture, Maharishi International University Fairfield, Iowa, USA 52556 massemaj@gmail.com text Zootaxa 2026 2026-01-14 5748 3 354 368 https://doi.org/10.11646/zootaxa.5748.3.2 journal article 314832 10.11646/zootaxa.5748.3.2 9963998f-ed92-4ba2-baa7-24dde3667d7f 1175-5326 19168509 7059F8F0-0CBB-49AD-B60B-105DBA00EDD5

TABLE 1. Comparison of species of the Philippine sangirensis group and P. sangirensis of Indonesia. Setal counts are given as a pair of numbers separated by a comma, the first of the pair being the count in segment vii, and the second being the number in xx. Setal gaps are represented as D for dorsal gap, either present (+D) or absent (–D), and likewise V for ventral. Reproductive pore spacings are given as the estimated fraction of body circumference. Typhlosole presence accompanied by the segment number of its origin if not in XXVII. Septal presence and absence are indicated by + or –, the first of each pair being for septum 8/9. For bursal pads, those found on the floor of the bursa are represented by a number followed by F (e.g. 4F), and those attached to the roof of the bursa are represented by a number followed by R (e.g. 4R). Missing data are shown by a question mark (?). Data from Aspe & James (2014; 2016; 2017), Hong & James (2008; 2011; 2021), – James (2004b), and Michaelsen (1896). *individuals with spermathecae; some are athecal.

	quincunxia	diesmosi	rugosa	monoporata	vicinipora	baungon-ensis	pauci-setosa	alba	rubida	virgata	sangirensis	apoensis	baracatan-ensis	camiguin-ensis	floresi	sibucal-ensis	solisi	timpoong-ensis	malindang-ensis	misamis-ensis	wati	nunezae	tigris	longi-prostata	maculo-dorsalis	immanis	lago	boniao	mariae	lagunaensis	vergrandis	concepcionensis	riparian-ensis	bontoc-ensis	banahaw-ensis sp. nov.	Pheretima sp 2 aurora	sancta sp. nov.	taceae sp. nov.	apolinarioi sp. nov.	madii sp. nov.
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setae vi, xxv	18, 25	28, 38	24, 32	30, 34	52, 40	38, 50	22, 28	38, 36	34, 36	76, 80	?	38–48, 50–58	28, 38	32, 42	37, 45	70, 90	52, 40	29–43, 50–72	44–48, 50–58	42–51, 43–48	60–70, 52–60	46, 51	38, 48–60	36–40, 38–48	74, 64–74	62–68, 64–68	49, 54	60–78, 62–68	32, 32	46, 52	37–42; 50–54	40–50; 56–68	42, 52	40, 62	46, 64	?	28, 38	31, 36	39, 45	52, 56
setal gaps D, V	Irreg.	+D, –V	+D, –V	+D, –V	+D, –V	+D, –V	+D, –V	+D, –V	+D, –V	–D, –V	+D,?V	+D,–V	+D, –V	+D, –V	+D, –V	+D, –V	+D, –V	+D, –V	–D, +V	+D, –V	+D, –V	+D, –V	+D, –V	+D, –V	+D, –V	–D, –V	+D, –V	+D, –V	+D, –V	+D, –V	+D, +V	+D, +V	+D, +V	+D, +V	+D, +V	-D, –V	–D, +V	+D, –V	+D, +V	+D, +V
male pore setae	0–2	0	0	0	0	8	2	0	4	4–5	6	4–5	4–5	1–4	5	9	3–5	2–5	0	6–7	0	9	0	0–5	2–4	5	0–2	0	1	0	–	2	5–7	4–6	4–7	5	6–7	1	0	0
male pore spacing	0.13	0.15	0.15	0.05	0.04	0.19	0.13	0.10	0.12	0.09	0.17–0.25	0.15	0.22	0.17	0.20	0.17	0.15	0.23	0.11	0.23	0.08	0.22	0.14	0.16	0.13	0.14	0.15	0.03	0.09	0.10	–	0.14	0.11	0.10	0.09–0.13	0.23	0.16	0.13	0.12	0.04
sp. pore spacing	0.16	0.23	0.20	0	0.08	0.38	0.16	0.13	0.20	0.11	0.25–0.28	0.16*	0.32	0.17	0.21	0.21	0.18	0.20	0.16	0.30	0.17	0.28	0.13	0.16	0.09	0.12	0.18–0.24	0.14	0.04	0.03	–	–	0.07	0.09	0.09–0.13	0.14	0.17–0.22	0.12	0	0.04
dorsal pores	12/13	12/13	11/12	12/13	13/14	13/14	11/12	13/14	11/12	12/13	?	12/13	12/13	12/13	12/13	12/13	12/13	12/13	12/13	12/13	12/13	12/13	12/13	11/12	12/13	12/13	12/13	?	12/13	12/13	13/14	12/13	12/13	12/13	4/5	11/12	12/13	5/6	11/12	12/13
dorsal pigment	–	brown	brown	brown	brown	brown	–	–	red-brown	brown stripes	?	violet brown	brown	red-brown	red- brown	gray brown	brown	violet brown	dark brown	black	brown	gray-brown	red - purple	yellowbrown	red	dark violet	dark brown	violet brown	dark violet	brown	light brown	dark brown	violet brown	brown	brown,	red	brown	brown	brown	brown
equatorial pigment	none	+	+	+	+	+	none	none	+	none	?	none	+	none	+	none	+	none	none	none	+	none	none	none	none	none	none	none	+	+	present	present	+	none	none	none	+	none	+	none
length mm	65–86	128	70	60–70	50	107	115	49	72	290–360	?	160–350	100	63–122	48–67	>165	68–86	145–225	60–80	55–65	67–75	>116	230–280	37–41	226–235	365	233–315	101–133	55	40	35–75	>89	75–105	68–83	130	90	65	80	33	54–60
intestinal origin	½xvi	xvii	xvii	xvii	xvi	xv	½xvi	xvi	xvi	xvi	?	xv	xv	xv	xv or xvi	xiv	xv	xv or xvi	xvi	xv	xv	xv	xvi	xv	xvii	xvi	xiv	xiv	xvi	xvi	xv	xvi	xvi	xvi	xvi	xvii	xvi	xvi	xvi	xvii
typhlosole	–	+	+	+	+	+	–	+	+	+	?	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+ 33	+ 33	+
esoph. lamellae	+	+	+	+	+	+	+	+	+	+	?	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	+	?	?	–	–	?	?	+	?	+	+	?	?
intestinal vessels	?	?	?	18–20	26	32	34	24	26	32–42	?	32–38	28	26–30	35	38	29	31–44	30	32	34–38	20–23	56–58	20	50–54	28–32	36–38	33–45	34–36	28–30	24–28	30–35	?	28–30	30	?	?	?	?	?
hearts	x–xiii	x– xiii	x–xiii	x–xiii	xi–xiii	x–xiii	x–xiii	x–xiii	x–xiii	x–xiii	?	x–xiii	x– xiii	x–xiii	x–xiii	x–xiii	xi–xiii	x–xiii	xi–xiii	x–xiii	x–xiii	xi–xiii	x–xiii	x–xiii	x–xiii	x–xiii	x–xiii	xi–xiii	x–xiii	x–xiii	x–xiii	x–xiii	x–xiii	x–xiii	x–xiii	x–xiii	x– xiii	x–xiii	x–xiii	x–xiii
septa 8/9/10	+, +	+, +	+, +	+, +	+, –	+, –	+, +	–, –	+, +	+, –	?	+, –	+, –	+, –	+, –	+, –	+, –	–, –	–, –	+, +	–, +	+, –	+, –	–, –	+, –	+, –	+, –	+, –	–, –	–, –	–,–	–,–	–, –	–, –	–, –	–, –	–, –	–, –	–, –	+, +
bithecate vs mono	bi	bi	bi	mono	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	bi	mono	mono	bi	bi	bi	bi	bi	bi	mono	bi
thecal segm.	vii	vii	vii	vii	viii	vii	vii	vii	vii	viii	?	viii*	viii	viii	viii	vii	viii	viii	viii	vii	viii	vii	viii	vii	viii	viii	vii	viii	vii	vii	vii	viii	vii	vii	vii	viii	vii	vii	viii	vii
penes	–	–	+	+	–	–	+	+	+	+	+	–	–	+	–	–	–	+	+	+	+	+	–	+	+	–	+	–	+	+	+	+	+	+	+	+	+	+	+	+
bursal pads	4F	2F	3F	0	2R	2R	3F	0	0	5F, 4R	?	1R	3R	1R	2R	1R	2R 1F	2R	1R, 2F	1F	1R	2R, 2F	5F	0	2R	2F	2R,1F	1R	0	0	0	0	ring of wedges	4F wedges	5F, 1R	2F oval pads	4F, 1R	ring F	5–6F wedges	4–5F wedges