<document tableId="C8F78EF5FF98A134FF2DFB25949EC373" sourceDocId="E8181713FF9DA131FFBAFFD19662C642" ID-DOI="http://doi.org/10.5281/zenodo.16782663" ID-Zenodo-Dep="16782663" captionStart="TABLE 1" checkinTime="1675762064165" checkinUser="tatiana" pageId="5" pageNumber="506" updateTime="1754680665236" updateUser="ExternalLinkService"><mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
    <mods:titleInfo>
        <mods:title>Omorgus (Omorgus) khandesh (Coleoptera: Scarabaeoidea: Trogidae), a new species from India, along with an annotated catalogue of the Oriental and Palearctic species of the subgenus Omorgus Erichson, 1847</mods:title>
    </mods:titleInfo>
    <mods:name type="personal">
        <mods:role>
            <mods:roleTerm>Author</mods:roleTerm>
        </mods:role>
        <mods:namePart>Strümpher, Werner P.</mods:namePart>
        <mods:affiliation>Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa.</mods:affiliation>
    </mods:name>
    <mods:name type="personal">
        <mods:role>
            <mods:roleTerm>Author</mods:roleTerm>
        </mods:role>
        <mods:namePart>Kalawate, Aparna S.</mods:namePart>
        <mods:nameIdentifier type="ORCID">0000-0001-6595-6749</mods:nameIdentifier>
        <mods:affiliation>Zoological Survey of India, Western Regional Centre, Vidhya Nagar, Sector- 29, P. C. N. T. (PO), Rawet Road, Akurdi, Pune, Maharashtra 411044, India. aparna _ ent @ yahoo. co. in; https: // orcid. org / 0000 - 0001 - 6595 - 6749</mods:affiliation>
        <mods:nameIdentifier type="email">aparna_ent@yahoo.co.in</mods:nameIdentifier>
    </mods:name>
    <mods:typeOfResource>text</mods:typeOfResource>
    <mods:relatedItem type="host">
        <mods:titleInfo>
            <mods:title>Zootaxa</mods:title>
        </mods:titleInfo>
        <mods:part>
            <mods:date>2023</mods:date>
            <mods:detail type="pubDate">
                <mods:number>2023-02-02</mods:number>
            </mods:detail>
            <mods:detail type="volume">
                <mods:number>5231</mods:number>
            </mods:detail>
            <mods:detail type="issue">
                <mods:number>5</mods:number>
            </mods:detail>
            <mods:extent unit="page">
                <mods:start>501</mods:start>
                <mods:end>522</mods:end>
            </mods:extent>
        </mods:part>
    </mods:relatedItem>
    <mods:location>
        <mods:url>http://dx.doi.org/10.11646/zootaxa.5231.5.1</mods:url>
    </mods:location>
    <mods:classification>journal article</mods:classification>
    <mods:identifier type="CLB-Dataset">53658</mods:identifier>
    <mods:identifier type="DOI">10.11646/zootaxa.5231.5.1</mods:identifier>
    <mods:identifier type="GBIF-Dataset">30c14449-ed8b-4f00-b877-9ef2c72adb08</mods:identifier>
    <mods:identifier type="ISSN">1175-5326</mods:identifier>
    <mods:identifier type="Zenodo-Dep">7609508</mods:identifier>
    <mods:identifier type="ZooBank">22143C8D-9639-4B87-8D8C-C3BB421534A1</mods:identifier>
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<table id="C8F78EF5FF98A134FF2DFB25949EC373"><caption pageId="5" pageNumber="506" startId="5.[151,239,1268,1292]"><emphasis bold="true" box="[151,267,1268,1292]" pageId="5" pageNumber="506">TABLE 1.</emphasis> Summary of morphological distinction between <taxonomicName _authorityPlausibility="0.78287446" _evidence="statusLabel" _step="newSpecies" authority="Strumpher &amp; Kalawate" authorityName="Strumpher &amp; Kalawate, new" authorityYear="2023" box="[792,1436,1268,1293]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="khandesh" status="new" subGenus="Omorgus"><emphasis box="[792,890,1269,1292]" italics="true" pageId="5" pageNumber="506">Omorgus</emphasis> (<emphasis box="[906,1004,1269,1292]" italics="true" pageId="5" pageNumber="506">Omorgus</emphasis>) <emphasis box="[1021,1122,1268,1292]" italics="true" pageId="5" pageNumber="506">khandesh</emphasis> Strümpher &amp; Kalawate, new</taxonomicName> <emphasis bold="true" box="[151,230,1304,1328]" pageId="5" pageNumber="506">species</emphasis> and <taxonomicName _authorityPlausibility="0.99930793" _evidence="catalogs" _step="species1" authority="(Haaf, 1954)" baseAuthorityName="Haaf" baseAuthorityYear="1954" box="[282,758,1304,1329]" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="rimulosus" subGenus="Omorgus"><emphasis box="[282,380,1305,1328]" italics="true" pageId="5" pageNumber="506">Omorgus</emphasis> (<emphasis box="[395,493,1305,1328]" italics="true" pageId="5" pageNumber="506">Omorgus</emphasis>) <emphasis box="[509,614,1304,1328]" italics="true" pageId="5" pageNumber="506">rimulosus</emphasis> (Haaf, 1954)</taxonomicName>.</caption><thead><tr id="22B8DC3FFF985ECEFF1CFA9093ECC33F"><th box="[166,468,1345,1405]" id="6169B543FF985ECEFF1CFA9097B6C33F" pageId="5" pageNumber="506">Characters</th><th box="[518,925,1345,1405]" id="6169B543FF985ECEFDBCFA9095FFC33F" pageId="5" pageNumber="506"><taxonomicName _authorityPlausibility="0.99946713" _evidence="statusLabelOccurrence" _step="newLabel2" authority="Strumpher" authorityName="Strumpher &amp; Kalawate, new" authorityYear="2023" box="[518,875,1345,1369]" class="Insecta" family="Trogidae" genus="Omorgus" id="5B88A5FEFF98A134FDBCFA909509C31B" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="khandesh" subGenus="Omorgus"><emphasis box="[518,611,1345,1368]" id="AEFC026FFF98A134FDBCFA909401C31A" italics="true" pageId="5" pageNumber="506">Omorgus</emphasis> (<emphasis box="[626,651,1345,1368]" id="AEFC026FFF98A134FDC8FA9094E9C31A" italics="true" pageId="5" pageNumber="506">O.</emphasis>) <emphasis box="[666,762,1345,1369]" id="AEFC026FFF98A134FD20FA909498C31B" italics="true" pageId="5" pageNumber="506">khandesh</emphasis> Strümpher</taxonomicName> &amp; Kalawate <emphasis bold="true" box="[620,745,1380,1405]" id="AEFC026FFF98A134FDD6FAB4948BC33E" pageId="5" pageNumber="506"><taxonomicNameLabel box="[620,745,1380,1405]" id="B5CFBF14FF98A134FDD6FAB4948BC33E" pageId="5" pageNumber="506" rank="species">new species</taxonomicNameLabel></emphasis></th><th box="[970,1422,1345,1405]" id="6169B543FF985ECEFC70FA9093ECC33F" pageId="5" pageNumber="506"><taxonomicName _authorityPlausibility="0.7827949" _evidence="statusLabel" _step="species1" authority="(Haaf, 1957) Kalawate" authorityName="Kalawate" authorityYear="2023" baseAuthorityName="Haaf" baseAuthorityYear="1957" box="[970,1353,1345,1369]" class="Insecta" family="Trogidae" genus="Omorgus" id="5B88A5FEFF98A134FC70FA90932BC31B" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="rimulosus" status="sp. nov." subGenus="Omorgus"><emphasis box="[970,1063,1345,1368]" id="AEFC026FFF98A134FC70FA909245C31A" italics="true" pageId="5" pageNumber="506">Omorgus</emphasis> (<emphasis box="[1078,1103,1345,1368]" id="AEFC026FFF98A134FB8CFA90922DC31A" italics="true" pageId="5" pageNumber="506">O.</emphasis>) <emphasis box="[1118,1218,1345,1369]" id="AEFC026FFF98A134FBE4FA9092A0C31B" italics="true" pageId="5" pageNumber="506">rimulosus</emphasis> (<bibRefCitation author="Haaf, E." box="[1231,1346,1345,1369]" id="F819A38CFF98A134FB75FA909320C31B" pageId="5" pageNumber="506" pagination="692 - 695" refId="ref9103" refString="Haaf, E. (1957) Drei neue Trox - Arten (Col. Scarab). Entomologische Arbeiten aus dem Museum G. Frey, 8, 692 - 695." type="journal article" year="1957">Haaf, 1957</bibRefCitation>)</taxonomicName></th></tr></thead><tbody><tr id="22B8DC3FFF985ECEFF1CFA5C93ECC38B"><th box="[166,468,1421,1481]" id="6169B543FF985ECEFF1CFA5C97B6C38B" pageId="5" pageNumber="506">Head</th><td box="[518,925,1421,1481]" id="6169B543FF985ECEFDBCFA5C95FFC38B" pageId="5" pageNumber="506">Median tubercle on clypeus low, indistinct</td><td box="[970,1422,1421,1481]" id="6169B543FF985ECEFC70FA5C93ECC38B" pageId="5" pageNumber="506">Median tubercle on clypeus distinct</td></tr><tr id="22B8DC3FFF985ECEFF1CFA0893ECC057"><th box="[166,468,1497,1557]" id="6169B543FF985ECEFF1CFA0897B6C057" pageId="5" pageNumber="506">Elytra: Intercostal punctures</th><td box="[518,925,1497,1557]" id="6169B543FF985ECEFDBCFA0895FFC057" pageId="5" pageNumber="506">Distinct, more closely spaced</td><td box="[970,1422,1497,1557]" id="6169B543FF985ECEFC70FA0893ECC057" pageId="5" pageNumber="506">Less pronounced, shallow more widely spaced</td></tr><tr id="22B8DC3FFF985ECEFF1CF9F493ECC07C"><th box="[166,468,1573,1598]" id="6169B543FF985ECEFF1CF9F497B6C07C" pageId="5" pageNumber="506">Elytra: even numbered costae</th><td box="[518,925,1573,1598]" id="6169B543FF985ECEFDBCF9F495FFC07C" pageId="5" pageNumber="506">Smaller, more closely spaced out</td><td box="[970,1422,1573,1598]" id="6169B543FF985ECEFC70F9F493ECC07C" pageId="5" pageNumber="506">Large, widely spaced out</td></tr><tr id="22B8DC3FFF985ECEFF1CF9A793ECC094"><th box="[166,468,1654,1790]" id="6169B543FF985ECEFF1CF9A797B6C0BC" pageId="5" pageNumber="506" rowspan="2">Parameres</th><td box="[518,925,1654,1750]" id="6169B543FF985ECEFDBCF9A795FFC094" pageId="5" pageNumber="506">Apex of each feebly sclerotised without distinct membranous projection (<figureCitation box="[526,611,1726,1750]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 11–16. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 11–13. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 14–16. a–j, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" id="04B3C2F8FF98A134FDB4F96F9401C094" pageId="5" pageNumber="506">Fig. 12a</figureCitation>)</td><td box="[970,1422,1654,1750]" id="6169B543FF985ECEFC70F9A793ECC094" pageId="5" pageNumber="506">Apex of each feebly sclerotised with distinct membranous projection (<figureCitation box="[1224,1308,1690,1714]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 11–16. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 11–13. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 14–16. a–j, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" id="04B3C2F8FF98A134FB72F94B937EC0F0" pageId="5" pageNumber="506">Fig. 15e</figureCitation>)</td></tr><tr id="22B8DC3FFF985ECEFF1CF93793ECC0BC"><td box="[518,925,1766,1790]" id="6169B543FF985ECEFDBCF93795FFC0BC" pageId="5" pageNumber="506">Narrower in lateral profile (<figureCitation box="[799,885,1766,1790]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 11–16. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 11–13. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 14–16. a–j, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" id="04B3C2F8FF98A134FCA5F9379517C0BC" pageId="5" pageNumber="506">Fig. 12b</figureCitation>)</td><td box="[970,1422,1766,1790]" id="6169B543FF985ECEFC70F93793ECC0BC" pageId="5" pageNumber="506">Broader in lateral profile (<figureCitation box="[1235,1318,1766,1790]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 11–16. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 11–13. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 14–16. a–j, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" id="04B3C2F8FF98A134FB69F9379344C0BC" pageId="5" pageNumber="506">Fig. 15f</figureCitation>)</td></tr><tr id="22B8DC3FFF985ECEFF1CF8DF93ECC108"><th box="[166,468,1806,1907]" id="6169B543FF985ECEFF1CF8DF97B6C131" pageId="5" pageNumber="506" rowspan="2">Phallus</th><td box="[518,925,1806,1866]" id="6169B543FF985ECEFDBCF8DF95FFC108" pageId="5" pageNumber="506">Anteromedially deeply concave, lateral protuberances prominent (<figureCitation box="[785,871,1842,1866]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 11–16. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 11–13. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 14–16. a–j, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" id="04B3C2F8FF98A134FCABF8E39505C108" pageId="5" pageNumber="506">Fig. 12d</figureCitation>)</td><td box="[970,1422,1806,1866]" id="6169B543FF985ECEFC70F8DF93ECC108" pageId="5" pageNumber="506">Anteromedially shallowly concave, lateral protuberances low (<figureCitation box="[1170,1256,1842,1866]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 11–16. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 11–13. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 14–16. a–j, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" id="04B3C2F8FF98A134FB28F8E3928AC108" pageId="5" pageNumber="506">Fig. 15h</figureCitation>)</td></tr><tr id="22B8DC3FFF985ECEFF1CF88A93ECC131"><td box="[518,925,1883,1907]" id="6169B543FF985ECEFDBCF88A95FFC131" pageId="5" pageNumber="506">Apex pointed; margin straight (<figureCitation box="[836,916,1883,1907]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 11–16. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 11–13. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 14–16. a–j, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" id="04B3C2F8FF98A134FCFEF88A95F6C131" pageId="5" pageNumber="506">Fig. 13i</figureCitation>)</td><td box="[970,1422,1883,1907]" id="6169B543FF985ECEFC70F88A93ECC131" pageId="5" pageNumber="506">Apex rounded; margin recurved (<figureCitation box="[1308,1389,1883,1907]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 11–16. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 11–13. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 14–16. a–j, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" id="04B3C2F8FF98A134FAA6F88A930FC131" pageId="5" pageNumber="506">Fig. 16j</figureCitation>)</td></tr><tr id="22B8DC3FFF985ECEFF1CF85293ECC1D9"><th box="[166,468,1923,1947]" id="6169B543FF985ECEFF1CF85297B6C1D9" pageId="5" pageNumber="506">Phallobasis (in lateral profile)</th><td box="[518,925,1923,1947]" id="6169B543FF985ECEFDBCF85295FFC1D9" pageId="5" pageNumber="506">Small (<figureCitation box="[591,676,1923,1947]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 11–16. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 11–13. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 14–16. a–j, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" id="04B3C2F8FF98A134FDF5F85294C6C1D9" pageId="5" pageNumber="506">Fig. 12c</figureCitation>)</td><td box="[970,1422,1923,1947]" id="6169B543FF985ECEFC70F85293ECC1D9" pageId="5" pageNumber="506">Larger (<figureCitation box="[1051,1137,1923,1947]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[283,1306,194,1920]" captionTargetId="figure-22@8.[267,1321,181,1920]" captionTargetPageId="8" captionText="FIGURES 11–16. Male genitalia in dorsal (11, 14) and lateral (12, 15) views, apex of median lobe (13, 16). Omorgus (Omorgus) khandesh Strümpher &amp; Kalawate, new species, holotype (ZSI-WRC), 11–13. Omorgus (Omorgus) rimulosus (Haaf, 1957), holotype (NHMB), 14–16. a–j, for explanation refer to Table 1." figureDoi="http://doi.org/10.5281/zenodo.7609522" httpUri="https://zenodo.org/record/7609522/files/figure.png" id="04B3C2F8FF98A134FBA1F8529213C1D9" pageId="5" pageNumber="506">Fig. <quantity box="[1098,1137,1923,1947]" id="5B707398FF98A134FBF0F8529213C1D9" metricMagnitude="-2" metricUnit="kg" metricValue="1.5" pageId="5" pageNumber="506" unit="g" value="15.0">15g</quantity></figureCitation>)</td></tr></tbody></table></document>