Review of Moluccan Rattus (Rodentia: Muridae) with Description of Four New Species Author Fabre, Pierre-Henri Institut des Sciences de l’Évolution de Montpellier, Université de Montpellier, CNRS, IRD, EPHE, 34095 Montpellier, France & Mammal Section, The Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom & Institut Universitaire de France (IUF), Paris, France & Division of Vertebrate Zoology (Mammalogy), American Museum of Natural History, Central Park West at 79 th St, New York, New York 10024 - 5192, United States of America Author Miguez, Roberto Portela Mammal Section, The Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Author Holden, Mary Ellen Division of Vertebrate Zoology (Mammalogy), American Museum of Natural History, Central Park West at 79 th St, New York, New York 10024 - 5192, United States of America Author Fitriana, Yuli S. Museum Zoologicum Bogoriense, Research Centre for Biosystematics and Evolution, National Research and Innovation Agency (BRIN), Jln. Raya Jakarta-Bogor Km. 46, Cibinong 16911, Indonesia Author Semiadi, Gono Center for Applied Zoological Research, National Research and Innovation Agency (BRIN), Jln. Raya Jakarta-Bogor Km. 46 Cibinong, 16911, Indonesia Author Musser, Guy G. Division of Vertebrate Zoology (Mammalogy), American Museum of Natural History, Central Park West at 79 th St, New York, New York 10024 - 5192, United States of America Author Helgen, Kristofer M. Australian Museum Research Institute, Australian Museum, 1 William St, Sydney NSW 2010, Australia & Australian Research Council Centre for Australian Biodiversity and Heritage, University of New South Wales, Sydney, New South Wales, Australia † Guy G. Musser, 1936 - 2019, deceased. text Records of the Australian Museum 2023 2023-12-13 75 5 673 718 journal article 10.3853/j.2201-4349.75.2023.1783 66680ea5-7dbb-4f08-b8a6-ac488a357e64 2201-4349 16780477

Table 2. Selected external measurements (mm) and body weight (g) of Rattus species from Maluku. Mean, range (in brackets) and sample size (after the brackets) are reported in each case. All specimens are adult except for the two specimens of Rattus obiensis sp. nov.

Taxa / islands	Sex	HB	TL	HF	E	WT	TL/HB (%)	HF/HB (%)
Seram Rattus ceramicus	♂ ♀	130.0 [125–135] 2 118	135 [130–140] 2 126	29.5 [29–30] 2 28	18 [18–18] 2 17	66.5 —	104 107	23 24
Rattus feliceus	♂ ♀	197.3 [167–225] 3 243.5 [200–287] 2	180 [175–185] 3 165 [165–165] 1	45.0 [44–46] 3 41.5 [40–43] 2	22 [22–22] 3 21 [20–22] 2	120 306 [272–345] 3	91 68	21 17
Taliabu Rattus elaphinus	♂	189.1 [161–215] 11	176 [154–188] 11	35.8 [34–38] 11	20.1 [18–22] 11	—	93	19
Rattus feileri sp. nov. Rattus taliabuensis sp. nov.	♀ ♂ ♂	173.6 [128–212] 17 176 234	172.2 [136–201] 17 237 165	34.9 [32–38] 17 40 43	19.5 [17–22] 17 17 24	— — —	99 134 71	20 23 18
Mangole Rattus elaphinus	♂	186.5 [173–200] 3	156 [156–156] 3	33.0 [33.0–34.6] 3	21.5 [19.6–22.5] 3	154.7 [123–197] 3	101	18
Morotai Rattus morotaiensis	♂ ♀	190.8 [167–221] 4 161.0 [148–184] 3	209.2 [20–216] 3 188 [165–213] 3	39.2 [38–41] 4 34.4 [30–35] 3	— 17	— 86	110 117	21 21
Halmahera
Rattus halmaheraensis sp. nov.	♂ ♀	163.6 [161–215] 17 151.2 [119.4–175] 19	186.8 [154–188] 17 184.2 [158–221] 18	34.0 [34–38] 17 33.3 [30.7–37.2] 19	19.3 [18–22] 17 19.9 [18.2–22.3] 19	112.5 [49–159] 17 103 [51–146] 17	114 122	21 22
Bacan Rattus halmaheraensis sp. nov.	♂	200	224	39.5	18.7	158	112	20
Obi Rattus obiensis sp. nov.	juv.♀	119.0 [116–122] 2	157.0 [157–157] 2	27.8 [27.3–28.3] 2	16.8 [16.7–16.9] 2	37.5 [36–39] 2	132	23