Systematics of African rough-scaled lizards, with description of two new species from eastern Angola (Squamata: Lacertidae: Ichnotropis Peters, 1854) Author Conradie, Werner 0000-0003-0805-9683 Port Elizabeth Museum (Bayworld), Beach Road, Humewood, Gqeberha, 6013, South Africa & Department of Conservation Management, Natural Resource Science and Management Cluster, Faculty of Science, George Campus, Nelson Mandela University, George, 6529, South Africa & National Geographic Okavango Wilderness Project, Wild Bird Trust, Santon, 2196, South Africa Author Keates, Chad 0000-0003-3125-9536 National Geographic Okavango Wilderness Project, Wild Bird Trust, Santon, 2196, South Africa & South African Institute for Aquatic Biodiversity, Makhanda, 6140, South Africa & Ft. Lauderdale Research and Education Center, University of Florida, Davie, FL 33314, USA Author Greenbaum, Eli 0000-0003-1674-1316 Department of Biological Sciences, University of Texas at El Paso, El Paso, TX 79968, USA Author Lobón-Rovira, Javier 0000-0003-4380-9427 CIBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, InBIO Laboratorio Associado, Campus de Vairao, Universidade do Porto, 4485 - 661 Vairao, Portugal & BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairao, 4485 - 661 Vairao, Portugal Author Tolley, Krystal A. 0000-0002-7778-1963 Department of Zoology, University of Johannesburg, Auckland Park, Johannesburg, 2006, South Africa Author Benito, Max https://orcid.org/0009-0000-6307-3053 Universidad Internacional Menéndez Pelayo, Calle de Isaac Peral 23, 28040 Madrid, Spain Author Vaz Pinto, Pedro CIBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, InBIO Laboratorio Associado, Campus de Vairao, Universidade do Porto, 4485 - 661 Vairao, Portugal & BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairao, 4485 - 661 Vairao, Portugal & Fundação Kissama, Rua 60 Casa 560, Lar do Patriota, Luanda, Angola Author van Breda, Reuben V. Unit for Environmental Sciences and Management, North-West University, Potchefstroom, 2531, South Africa Author Verburgt, Luke National Geographic Okavango Wilderness Project, Wild Bird Trust, Santon, 2196, South Africa & Enviro-Insight CC, Unit 8 Oppidraai Office Park, Pretoria, 0050, South Africa & Department of Zoology and Entomology, University of Pretoria, Pretoria, 0001, South Africa text Vertebrate Zoology 2025 2025-11-21 75 627 672 journal article 313199 10.3897/vz.75.e167366 4a5fd0e1-5ec9-470a-8457-788a0110566d F811EE38-D26A-4C49-A863-D2800F54BA7B

Table 6. Summary of morphological data for the Ichnotropis capensis group. Measurements are in millimetres (mm). Values are given as a range with mean ± standard deviation in parenthesis. Juveniles were excluded from the measurements, but were included in the scalation data. For abbreviations, see the Materials and Methods section. n = sample size.

Characters	I. capensis sensu lato	I. longicorpa sp. nov.
	n = 256	n = 12
SVL	40.0–67.8 (54.8 ± 5.11)	62.7–71.2 (66.4 ± 2.37)
TAIL	69.5–149.0 (110.4 ± 17.89)	117–160 (137.5 ± 15.94)
HL	10.1–14.7 (12.7 ± 0.97)	13.1–17.2 (15.1 ± 1.16)
HW	5.6–8.6 (7.2 ± 0.67)	7.8–9.4 (8.7 ± 0.46)
HH	4.5–7.5 (5.9 ± 0.62)	6.1–8.5 (7.4 ± 0.61)
ED	3.9–6.3 (4.5 ± 0.52)	2.4–5.9 (4.4 ± 0.98)
SE	4.0–7.0 (5.2 ± 0.77)	6.0–7.3 (6.9 ± 0.40)
LL	12.5–16.4 (14.2 ± 1.18)	14.4–18.7 (17.8 ± 1.27)
IL	18.5–31.6 (25.5 ± 3.29)	27.4–31.6 (28.7 ± 1.42)
FLL	5.0–8.2 (6.2 ± 0.70)	6.8–8.4 (7.5 ± 0.46)
HLL	7.4–13.2 (10.5 ± 1.25)	10.6–13.5 (12.3 ± 0.96)
TAIL / SVL	1.2–2.7 (2.0 ± 0.28)	1.9–2.4 (2.2 ± 0.22)
HL / SVL	0.2–0.3 (0.2 ± 0.01)	0.2–0.3 (0.2 ± 0.01)
ES / HL	0.4–0.5 (0.5 ± 0.02)	0.4–0.5 (0.4 ± 0.02)
HW / HL	0.2–0.7 (0.6 ± 0.06)	0.5–0.6 (0.6 ± 0.03)
MSR	25–42	34–41
LVSR	8 – 10	9–10
TVSR	20 – 31	25–31
SL	3 – 6 (mostly 4)	4–5 (mostly 4)
IL	5–8 (mostly 6)	6–7 (mostly 6)
SC	3–5	4
LUFT	16–26	19–24
Femoral pores	6–15	10–13