The genus Dryadaula Meyrick (Lepidoptera, Tineoidea, Dryadaulidae) in China, with descriptions of four new species and a world checklist Author Yang, Lin-Lin https://orcid.org/0000-0001-8540-5522 Institute of Plant Protection, Henan Academy of Agricultural Sciences, Zhengzhou 450002, China Author Li, Hou-Hun https://orcid.org/0000-0002-8953-3422 College of Life Sciences, Nankai University, Tianjin 300071, China lihouhun@nankai.edu.cn text ZooKeys 2021 2021-12-01 1074 61 81 http://dx.doi.org/10.3897/zookeys.1074.73067 journal article http://dx.doi.org/10.3897/zookeys.1074.73067 1313-2970-1074-61 A921393D58B048E18F101A9A163D8B91 52A4BA0B309A5C15944951BE15406EA4

Table 2. World checklist of the genus Dryadaula Meyrick, 1893.

	Species	Distribution	Depository of type
1	acrodisca (Meyrick, 1917): 79. (Choropleca)	Guyana	NHM (LT)
	TL: Guyana (Mallali).
2	amentata (Meyrick, 1919): 271. (Opsodoca)	Guyana	NHM (HT)
	TL: Guyana (Bartica). Figs: Clarke (1970: pl. 34, fig. 2 adult and male genitalia).
3	advena (Zimmerman, 1978): 326. (Choropleca)	United States	BMH (HT)
	TL: United States (Hawaii). Figs: Zimmerman (1978: fig. 156-A male genitalia; fig. 481-A adult).
4	anthracorma Meyrick, 1915: 369.	Australia	NHM (LT and PLT)
	TL: Australia (Victoria). Figs: Robinson and Nielsen (1993: fig. 63 adult; fig. 70 male genitalia; figs 71, 72 female genitalia); Robinson (2009: fig. 2 adult).
5	Dryadaula auriformis sp. nov.	China	NKU (HT and PT)
	TL: China (Hainan). Figs 1, 7.
6	boviceps (Walsingham, 1914): 366. (Choropleca)	Mexico	NHM (HT)
	TL: Mexico (Guerrero). Figs: Robinson (2009: fig. 4 adult).
7	brontoctypa (Meyrick, 1880): 259. (Ereunetis)	Australia	NHM (LT and PLT)
	TL: Australia (Sydney).
8	castanea Philpott, 1915: 201.	New Zealand	?
	TL: New Zealand (Bluff, Invercargill).
9	catorthota (Meyrick, 1917): 80. (Choropleca)	Guyana	NHM (LT and PLT)
	TL: Guyana (Mallali).
10	Dryadaula caucasica (Zagulajev, 1970): 662. (Archimeessia)	Azerbaijan, Poland, Russia, Sweden	ZIN (HT)
	TL: Azerbaijan (Artschevan). Figs: Zagulajev (1970: fig. 6 female genitalia); Zagulajev (1979: fig. 64 adult; fig. 65 female genitalia); Sachkov (1995: fig. 7 male genitalia); Jaworski et al. (2012: fig. 1 adult); Gaedike (2015: pl. 1, fig. 1 adult; drawings, male genitalia 1; drawings, female genitalia 1).
11	discatella (Walker, 1864): 1021. (Gelechia)	Brazil	NHM (HT)
	TL: Brazil.
12	Thermocrates epischista (Meyrick, 1936): 621. (Thermocrates)	China (Hong Kong), Japan	NHM (HT)
	TL: Japan (Kyushu). Figs: Robinson (1988: fig. 1 adult; fig. 2 abdominal pelt; figs 3, 4 male genitalia); Sakai (2013: fig. 3-12-13 adult).
13	epixantha (Turner, 1923): 184. (Erechthias)	Australia	ANIC
	TL: Australia (Queensland).
14	Dryadaula flavostriata sp. nov.	China	NKU (HT)
	TL: China (Guangxi). Figs 2, 8.
15	germana (Walsingham, 1914): 367. (Choropleca)	Mexico	NHM (HT)
	TL: Mexico (Guerrero).
16	Tinea glycinocoma (Meyrick, 1932): 120. (Tinea)	Ethiopia	NHM (LT and PLT)
	TL: Ethiopia. Figs: Gozmany and Vari (1973: fig. 6 male genitalia).
17	Dryadaula glycinopa Meyrick, 1893: 559.	Australia	NHM (LT)
	TL: Australia (New South Wales). Figs: Robinson and Nielsen (1993: fig. 64 adult; fig. 136 wing venation).
18	Dryadaula heindeli Gaedike & Scholz, 1998: 106.	Belgium, France, Germany, Italy, Netherlands, Norway, Spain, Switzerland	SDEI (HT and PT); coll. Scholz (PT); coll. Heindel (PT); coll. Sutter (PT)
	TL: Germany (Bayem). Figs: Gaedike and Scholz (1998: fig. 1 adult; figs 3-6 male genitalia; fig. 7 female genitalia; figs 10-15 larva, chaetotaxy and pupa); Gaedike (2015: pl. 1, fig. 5 adult; drawings, male genitalia 5; drawings, female genitalia 5).
19	hellenica (Gaedike, 1988): 331. (Archimeessia)	Greece	ZMUC (HT and PT); SDEI (PT).
	TL: Greece (Peloponnese). Figs: Gaedike (1988: figs 22-26 male genitalia); Gaedike (2015: pl. 1, fig. 7 adult; drawings, male genitalia 7; drawings, female genitalia 7).
20	Dryadaula hirtiglobosa sp. nov.	China	NKU (HT and PT)
	TL: China (Guangxi). Figs 3, 9.
21	Dryadaula irinae (Savenkov, 1989): 94. (Archimeessia)	Austria, Bulgaria, Latvia, Poland, Slovakia,	LMNH
	TL: Latvia. Figs: Savenkov (1989: figs. 1-3); Pastoralis et al. (2011: figs 1, 2 adults; fig. 3 male genitalia); Jaworski et al. (2014: fig. 5 adult; fig. 6 larva; fig. 7 larval shelter; fig. 8 pupal case); Gaedike (2015: pl. 1, fig. 3 adult; drawings, male genitalia 3; drawings, female genitalia 3).
22	isodisca (Meyrick, 1917): 80. (Choropleca)	Guyana	NHM (LT)
	TL: Guyana (Bartica, Mallali).
23	Dryadaula koreana Roh & Byun, 2020: 222	South Korea	SEL/ HNU (HT and PT)
	TL: South Korea (Jeollanam-do). Figs: Roh et al. (2020: figs. 1, 2 adult; fig. 3: wing venation; fig. 4 male genitalia; fig. 5 female genitalia).
24	Ditrigonophora marmoreipennis (Walsingham, 1897): 118. (Ditrigonophora)	Grenada	NHM (LT)
	TL: Grenada (Balthazar).
25	Chorocosma melanorma (Meyrick, 1893): 560. (Chorocosma)	Australia	NHM (HT)
	TL: Australia (Sydney).
26	mesosticha (Turner, 1923): 184. (Erechthias)	Australia	ANIC
	TL: Australia (Queensland).
27	Opsodoca metrodoxa (Meyrick, 1919): 271. (Opsodoca)	Guyana	NHM (LT)
	TL: Guyana (Bartica, Mallali). Figs: Clarke (1970: pl. 34, fig. 1 adult, wing venation and male genitalia).
28	minuta Gaedike, 2007: 160.	Greece, Turkey	ZMHB (HT), ZMUC (PT)
	TL: Turkey (Mugla). Figs: Gaedike (2007: fig. 1 adult; figs 12, 13 male genitalia; fig. 14 female genitalia); Gaedike (2015: pl. 1, fig. 4 adult; drawings, male genitalia 4; drawings, female genitalia 4).
29	multifurcata Gaedike, 2000: 358.	Russia	ZIN (HT and PT)
	TL: Russia (Primorskij kraj). Figs: Gaedike (2000: figs 5-7 male genitalia; fig. 8 female genitalia).
30	murenula (Meyrick, 1924): 65. (Choropleca)	Peru	NHM (LT and PLT)
	TL: Peru (Jurimaguas, Iquitos).
31	myrrhina Meyrick, 1905: 243.	New Zealand	NHM (HT)
	TL: New Zealand. Robinson (2009: fig. 3 adult).
32	napaea Meyrick, 1905: 244.	Australia	NHM (HT)
	TL: Australia (Tasmania).
33	Infurcitinea nedae (Gaedike, 1983): 125. (Infurcitinea)	Croatia, Cyprus, Greece, Turkey	coll. Baldizzone (HT and PT); SDEI (PT)
	TL: Greece. Figs: Baldizzone (1983: figs 3-7 male genitalia; fig. 8 female genitalia); Gaedike (2015: pl. 1, fig. 8 adult; drawings, male genitalia 8; drawings, female genitalia 8).
34	pactolia Meyrick, 1901: 577.	Denmark, France, Germany, Great Britain, New Zealand, Netherlands, Portugal, Switzerland.	NHM (LT)
	TL: New Zealand. Figs: Gaedike and Scholz (1998: fig. 2 adult; figs 8, 9 male genitalia); Gaedike (2015: pl. 1, fig. 6 adult; drawings, male genitalia 6; drawings, female genitalia 6).
35	panscia Meyrick, 1917: 81. (Choropleca)	Ecuador	NHM (LT)
	TL: Ecuador (Huigra).
36	placens Meyrick, 1920: 363. (Hectacma)	Australia	NHM (HT)
	TL: Australia (Queensland).
37	poecilta Walsingham, 1914: 366. (Choropleca)	Mexico	NHM (HT)
	TL: Mexico (Guerrero).
38	rhombifera Meyrick, 1917: 82. (Choropleca)	Guyana	NHM (HT)
	TL: Guyana (Mallali).
39	Dryadaula securiformis sp. nov.	China	NKU (HT and PT)
	TL: China (Hainan). Figs 4, 5, 6, 10 and 11.
40	selenophanes (Meyrick, 1880): 259. (Ereunetis)	Australia	NHM (LT)
	TL: Australia (Queensland).
41	sublimis (Meyrick, 1917): 81. (Choropleca)	Colombia	NHM (LT)
	TL: Colombia (La Crumbre).
42	Dryadaula terpsichorella (Busck, 1910): 134. (Cyane)	Fiji, Hawaii, Rapa.	NHM (HT)
	TL: Hawaii (Honolulu). Figs: Clarke (1971: pl. 28, figs. a, b adult; fig. 172 male genitalia, coremata and 8th segment; fig. 173 wing venation and female genitalia); Zimmerman (1978: fig. 157 head and wing venation; fig. 159 adult, male and female genitalia; fig. 160 abdomen and male genitalia); Robinson (2009: fig. 5 adult).
43	Dryadaula trapezoides (Meyrick, 1935): 579. (Tinea)	Japan	NHM (LT)
	TL: Japan (Tokyo). Figs: Sakai (2013: figs 3-12-14 adult; fig. Tin12 female genitalia).
44	tripudians (Meyrick, 1924: 65). (Choropleca)	Peru	NHM (LT)
	TL: Peru (Jurimaguas).
45	ussurica Gaedike, 2000: 358.	Russia	ZIN (HT and PT)
	TL: Russia (Primorskij kraj). Figs: Gaedike (2000: figs. 1-3 male genitalia; fig. 4 female genitalia).
46	Cyane visaliella (Chambers, 1873): 113. (Cyane)	Canada, United States	MCZ (ST)
	TL: United States (Kentucky). Figs: Zimmerman (1978: fig. 158 wing venation); Regier et al. (2015: fig. 6F adult; fig. 10 head, wing venation, male and female genitalia).
47	Archimeessia zinica (Zagulajev, 1970: 661). (Archimeesia)	Azerbaijan, Russia	ZIN (HT and PT); NHM (PT)
	TL: Azerbaijan. Figs: Zagulajev (1970: fig. 1 head; fig. 2 wing venation; fig. 3 legs; fig. 4 male genitalia; fig. 5 female genitalia); Gaedike (2015: pl. 1, fig. 2 adult; drawings, male genitalia 2; drawings, female genitalia 2).
48	zygodes Meyrick, 1918: 44. (Tinea)	South Africa	TM (HT)
	TL: South Africa (Natal). Figs: Janse (1968: pl. 69, fig. 3 adult, fig. 4 male genitalia; pl. 109: fig. 7 wing venation; pl. 111, fig. 16 labial palpi and maxillary palpi; pl. 113, fig. 9 male genitalia); Gozmany and Vari (1973: fig. 58 male genitalia)
49	zygoterma Meyrick, 1917: 82. (Choropleca)	Colombia, Ecuador	NHM (LT)
	TL: Colombia (La Crumbre).