Occurrence of the milk-eye catshark Apristurus nakayai (Carcharhiniformes: Pentanchidae) from the South China Sea Author Ng, Shing-Lai 0000-0001-6914-9100 Department of Environmental Biology and Fisheries Science, National Taiwan Ocean University, Keelung, Taiwan & terryzxq 1234567890 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6914 - 9100 terryzxq1234567890@gmail.com Author Liu, Kwang-Ming Center of Excellence for the Oceans, National Taiwan Ocean University, Keelung, Taiwan & George Chen Shark Research Center, National Taiwan Ocean University, Keelung, Taiwan & Institute of Marine Affairs and Resource Management, National Taiwan Ocean University, Keelung, Taiwan Author Joung, Shoou-Jeng Department of Environmental Biology and Fisheries Science, National Taiwan Ocean University, Keelung, Taiwan & George Chen Shark Research Center, National Taiwan Ocean University, Keelung, Taiwan text Zootaxa 2023 2023-02-16 5244 1 51 60 http://dx.doi.org/10.11646/zootaxa.5244.1.4 journal article 55174 10.11646/zootaxa.5244.1.4 949ff433-c0b9-444f-b3ea-8064ec10110f 1175-5326 7645767 A2EC8692-7E89-4C6F-A454-437770E099FC

TABLE 1 Morphometrical measurements of Apristurus nakayai. Data of the holotype and non-type (NTUM 11488) were from Iglésias (2012) and White et al. (2017), respectively.

	Non-types			Holotype
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Catalog number	NMMB-P36993 (n = 1)	EBFS 00138, NMMB- P36994–36997 (n = 4)	NTUM 11488 (n = 1)	MNHN 2003-1983
Sex	Mature female	Mature males	Mature male	Mature male	SD
Total length (mm)	408	415–451	559	659
Measurements (%TL)		Mean (Range)
Pre-D2 insertion	66.7	68.0 (66.7–69.9)	68.5	68.3	1.0
Pre-D2 origin	60.5	61.4 (59.9–63.0)	62.1	61.5	0.9
Pre-D1 insertion	53.2	53.2 (52.0–55.2)	52.6	52.2	1.0
Pre-D1 origin	49.8	48.5 (47.2–49.9)	47.4	47.6	1.0
Pre-P1 length	21.6	20.8 (19.5–21.6)	21.4	20.2	0.7
Pre-P2 length	38.0	37.1 (36.6–37.4)	37.2	37.2	0.4
Pre-vent length	43.0	41.0 (40.0–41.7)	41.9	40.4	0.9
Preanal length	52.2	51.5 (49.4–53.9)	47.4	51.6	1.9
Precaudal length	67.6	70.1 (68.6–71.8)	69.2	68.6	1.4
Pre-branchial length	18.4	17.2 (15.8–18.8)	16.9	15.9	1.0
Pre-spiracular length	13.8	13.4 (13.0–13.9)	11.9	12.1	0.7
Pre-orbital length	10.1	9.8 (9.2–10.5)	7.8	8.5	0.8
Pre-outer nostril length	4.9	4.6 (4.4–4.8)	4.5	4.2	0.2
Pre-inner nostril length	7.8	7.6 (7.3–8.1)	6.4	6.5	0.6
Pre-oral length	8.5	8.6 (8.3–8.9)	8.1	7.7	0.3
Head length	21.6	21.5 (21.2–22.1)	22.1	20.3	0.5
Head height	9.1	7.7 (7.1–8.3)	8.9	7.0	0.8
Head width at mouth corners	8.9	9.7 (9.5–9.9)	9.8	10.0	0.3
Head width max	9.2	10.2 (9.9–10.6)	10.1	10.6	0.4
Mouth width	6.4	6.6 (6.0–6.8)	8.3	6.8	0.6
Mouth length	3.8	4.3 (3.8–4.9)	2.6	2.4	0.8
Internarial width	3.6	3.4 (3.2–3.6)	3.4	3.3	0.1
Upper labial furrow length	3.3	3.6 (3.5–3.7)	2.9	3.1	0.2
Lower labial furrow length	2.5	1.8 (1.6–2.0)	2.1	2.1	0.3
Orbit length	3.8	3.7 (3.4–3.9)	3.3	3.2	0.2
Orbit height	1.5	1.1 (0.9–1.3)	0.9	1.1	0.2
Nostril length	4.0	3.6 (3.5–3.6)	3.2	3.0	0.3
Nostril -mouth space	1.8	1.8 (1.7–2.0)	1.7	1.7	0.1
Interorbital width	5.2	5.0 (4.7–5.5)	5.6	5.3	0.3
1st gill height	1.8	1.5 (1.2–1.7)	1.5	1.3	0.2
3rd gill height	2.0	2.0 (1.6–2.5)	1.4	1.7	0.3
5th gill height	1.5	1.7 (1.3–2.0)	1.5	1.5	0.2
D1-D2 space	6.7	8.9 (8.4–9.8)	9.5	9.4	0.9
D1-D2 origin	11.6	13.6 (12.4–15.0)	14.3	14.0	1.0
D1-D2 insertions	13.1	14.8 (13.4–15.9)	15.8	16.2	1.2
P1-P2 space	9.5	8.7 (8.1–9.6)	8.4	9.0	0.5
P1 tip to P2 origin	2.9	2.1 (1.8–2.8)	2.1	3.3	0.5
Catalog number	NMMB-P36993 (n = 1)	EBFS 00138, NMMB- P36994–36997 (n = 4)	NTUM 11488 (n = 1)	MNHN 2003-1983
Sex	Mature female	Mature males	Mature male	Mature male	SD
P1-P2 origins	15.7	15.5 (14.3–16.8)	16.7	16.4	0.9
P1-P2 insertions	18.2	15.5 (14.9–17.0)	16.1	15.4	1.1
P2-anal space	5.1	7.8 (6.3–8.6)	7.9	8.2	1.1
P2-anal origins	12.8	15.2 (14.8–15.9)	15.2	14.3	0.9
D1 length	7.2	7.4 (6.5–8.4)	7.2	7.2	0.5
D1 base length	4.5	5.1 (4.5–5.8)	5.0	4.8	0.4
D1 height	1.3	1.0 (0.8–1.2)	1.6	1.4	0.2
D1 free lobe length	2.9	2.7 (2.2–3.1)	2.5	2.7	0.3
D2 length	10.3	10.6 (9.0–11.6)	9.7	9.8	0.8
D2 base length	6.3	7.0 (6.0–7.9)	6.7	6.6	0.5
D2 height	2.4	2.1 (1.9–2.4)	2.8	2.2	0.3
D2 free lobe length	4.1	3.8 (3.3–4.2)	3.1	3.3	0.4
P1 base length	8.6	8.0 (7.3–8.7)	8.5	8.6	0.5
P1 anterior margin	14.0	12.7 (11.9–13.8)	13.1	12.4	0.8
P1 posterior margin	9.2	7.6 (7.1–8.8)	8.7	7.7	0.7
P1 inner margin	6.4	6.0 (3.6–6.8)	5.8	4.6	1.1
P1 width	8.8	7.9 (6.8–8.7)	–	9.0	0.7
P2 anterior margin	5.0	4.5 (4.0–4.7)	5.0	3.6	0.4
P2 length	11.8	9.5 (8.7–9.9)	10.5	9.0	0.9
P2 base length	9.4	7.1 (6.7–7.4)	8.3	7.6	0.8
P2 posterior margin	7.2	5.9 (5.4–6.2)	6.5	6.2	0.5
P2 inner margin	3.0	3.8 (3.5–4.5)	3.7	3.3	0.4
Anal base length (ceratotrichia)	15.2	16.1 (15.6–16.7)	16.7	17.5	0.7
Anal base length (muscle)	16.8	19.2 (17.9–21.2)	22.0	19.0	1.6
Anal anterior margin	7.7	7.7 (6.4–8.3)	6.6	8.0	0.7
Anal posterior margin	8.9	10.4 (9.1–11.6)	12.3	10.9	1.1
Anal height (muscle)	5.5	4.6 (3.7–5.2)	4.9	4.1	0.5
Anal inner margin	1.0	0.8 (0.7–0.9)	0.9	0.8	0.1
Caudal peduncle height	4.5	4.6 (4.4–4.8)	4.3	4.8	0.2
Caudal length	31.4	31.4 (30.2–32.7)	31.2	32.0	0.7
Caudal height	9.1	7.5 (7.0–8.2)	8.9	8.0	0.7
Caudal preventral margin	10.5	9.5 (7.9–10.7)	10.1	10.5	1.0
Caudal postventral margin	17.1	15.3 (14.2–16.7)	16.6	16.4	0.9
Caudal terminal lobe height	3.2	3.1 (2.9–3.4)	3.6	2.9	0.2
Caudal terminal lobe length	6.3	6.2 (6.0–6.5)	6.1	6.4	0.2
Clasper outer length	–	4.1 (3.4–4.5)	4.8	5.0	0.5
Clasper inner length	–	7.2 (6.7–7.7)	6.3	9.0	0.8