First tabulation and analysis of natural enemies of snail-killing flies (Diptera: Sciomyzidae), their position in the fly / mollusc ecosystem, and implications for use of sciomyzids in biological control Author Williams, C. D. Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; Author Knutson, L. Author Gordh, G. Plant Protection & Quarantine, APHIS USDA, Raleigh, NC, USA; Author Mc Donnell, R. J. Department of Crop and Soil Science, Oregon State University, Corvallis, OR, USA; Author Kula, R. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Gates, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Buffington, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Broad, G. Department of Entomology, Natural History Museum, London, UK & Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; text Journal of Natural History 2025 2025-01-30 59 5 - 8 331 396 https://doi.org/10.1080/00222933.2024.2443125 journal article 10.1080/00222933.2024.2443125 1464-5262 14798887

Table 3. (Continued).

Phygadeuon pumilis	HKT	Dictya sp.	(P-13) 13.iii-7.iv	New York	Knutson and Neff, unpub.
Phygadeuon sp. A, B, C	HKT	Tetanocera	“spring”	New York	Foote (1999)
		ferruginea
Phygadeuon sp.	LMW	Colobaea americana	(P) 26.iv-14.v/?	New York	Knutson and Bratt, unpub.
“	LMW	Tetanocera sp.	2.v/?	New York	Knutson, unpub.
“ ‘ Cryptine’ “ “	RWC A A A	Sciomyza varia Colobaea bifasciella Colobaea punctata Tetanocera	late iv-late v/early vi − − −	New York Denmark Denmark Denmark	Barnes (1990) Lundbeck (1923) Lundbeck (1923) Lundbeck (1923)
		ferruginea
Unidentified		Colobaea bifasciella	9.vi/10.vii	Denmark	Knutson and Bratt, unpub.
“		Anticheta	(P-3) 18.v/20.v, 6.vi	Denmark	Knutson (1966)
		brevipennis
“		Dictya steyskali		New York	Valley and Berg (1977)
“		Elgiva solicita	10.ii/21.ii	New York	Knutson and Berg (1964)
“ “		Elgiva solicita Renocera striata 16	“early spring/9.iv” “spring”	New York Ohio	Knutson and Berg (1964) Foote (1976)
“		Pherbina coryleti	(P-1) 29.iv/9.v	Greece	Knutson et al. (1975)
“		Tetanocera plebeja	(P-3) 7, 30.iv	New York, Idaho	Foote (1961)
“		Sciomyza simplex	(1L) 1.ix(ii.ix)/9.xi	Denmark	Knutson, unpub.
“		Sepedonea telson	3.v/16.v	Brazil	Abercrombie (1970)
“		Tetanocera annae	(P-1) 2.iv/14.iv	New York	Foote (1999)
“		Tetanocera	(P-2) 12.iii/24.iii	New York	Foote (1996a)
		fuscinervis
Unidentified (2 species)		Tetanocera	(P-23) 27.iii-2.iv/?	New York	Foote (1996b)
		rotundicornis
?		Sepedon fuscipennis	“summer”	New York	Eckblad and Berg (1972)
Hymenoptera
Braconidae
Aphaereta pallipes	CFWM	Atrichomelina	(P-”50”) 5.viii/?	Michigan	Foote et al. (1960)
		pubera
Aphaereta sp.	CFWM	Pherbellia subtilis (as	(P-1)		Bratt et al. (1969)
		obscura)
Panerema inops	RDE	Pherbellia sp.	(P-26?)/20.i-11.x	England,	Bratt et al. (1969)
				Denmark,
				Finland
Phaenocarpa antichaetae		Anticheta	(P-30) 21.iii/1-6.iv	New York	Fischer (1974), Knutson and Abercrombie
		melanosoma			(1977)
“		Anticheta	27.iv/8-11.v	New York	Fischer (1974), Knutson and Abercrombie
		melanosoma			(1977)