First tabulation and analysis of natural enemies of snail-killing flies (Diptera: Sciomyzidae), their position in the fly / mollusc ecosystem, and implications for use of sciomyzids in biological control Author Williams, C. D. Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; Author Knutson, L. Author Gordh, G. Plant Protection & Quarantine, APHIS USDA, Raleigh, NC, USA; Author Mc Donnell, R. J. Department of Crop and Soil Science, Oregon State University, Corvallis, OR, USA; Author Kula, R. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Gates, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Buffington, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Broad, G. Department of Entomology, Natural History Museum, London, UK & Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; text Journal of Natural History 2025 2025-01-30 59 5 - 8 331 396 https://doi.org/10.1080/00222933.2024.2443125 journal article 10.1080/00222933.2024.2443125 1464-5262 14798887

Table 3. (Continued).

Mastrus sp.?	WRMM	Anticheta	21.iii - 8.iv	New York	Knutson and Abercrombie (1977)
		melanosoma
“	WRMM	Anticheta	27.iv - 12.v	New York	Knutson and Abercrombie (1977)
		melanosoma
Orthizema n. sp.? Possible	RWC	Colobaea americana	(7) 27.iv - 18/21.v	New York	Knutson and Bratt, in prep.
misidentification
Phygadeuon sp. (misidentified	A	Sepedon sphegea	viii/?	Germany	Gercke (1876)
as ‘ cinctorius’)
Phygadeuon elegans	?	Tetanocera	(P-1) 17.x/xii	England	Disney (1964)
		ferruginea
Phygadeuon leucostigmus	HKT	Pherbina coryleti	(P-21) 7,9.iv/10.iv-18.v	Italy	Knutson et al. (1975)
Phygadeuon trichops	BL	Sepedon h.	(P-1) 7.ii / 19.ii	Spain	Knutson et al., 1967
		hispanica
Phygadeuon yonedai Phygadeuon sp. A, B	? WRMM	Sepedon aenescens Anticheta borealis	− −	Japan Ohio	Kusigamati (1986) Robinson and Foote (1978)
Phygadeuon sp. C	HKT	Pherbellia dorsata	(P-2) 27, 28.vi/?	Denmark	Bratt et al. (1969) and Bratt, unpub.
Phygadeuon sp. 1 nr. trichops	WRMM	Anticheta	28.iv/8,14.v	New York	Knutson and Abercrombie (1977)
		melanosoma
“	WRMM	Anticheta	14.iv/6.v	New York	Knutson and Abercrombie (1977)
		melanosoma
Phygadeuon sp. 2	WRMM	Anticheta	21.iii-2.iv	New York	Knutson and Abercrombie (1977)
		melanosoma
“	WRMM	Anticheta	14.iv/23.iv	New York	Knutson and Abercrombie (1977)
Phygadeuon sp. 7	WRMM	melanosoma Ditaeniella humilis	−	−	Bratt et al. (1969)
“	WRMM	Anticheta	21.iii, 27.iv -?	New York	Knutson and Abercrombie (1977)
Phygadeuon sp.	CFWM	melanosoma Atrichomelina	−	New York,	Foote et al. (1960)
“		pubera Anticheta testacea 15	−	Michigan California	Fisher and Orth (1964)
“	?	Sepedon pacifica		California	Fisher and Orth (1983)
“		Tetanocera	“spring”	New York	Foote (1999)
		ferruginea
Phygadeuontinae sp. Cylindricomorpha ’ sp.	RWC	Pherbellia cinerella Colobaea sp. ?	25.iv - 30.iv −	Ireland Denmark?	Gormally (1987b) Knutson, unpub.
Phygadeuon pumilis	HKT	Tetanocera	“spring”	New York	Foote (1999)
		ferruginea
Phygadeuon pumilis	HKT	Pherbellia sp.	(P-1) 8.iv/?	New York	Bratt et al. (1969) and Bratt, unpub.