First tabulation and analysis of natural enemies of snail-killing flies (Diptera: Sciomyzidae), their position in the fly / mollusc ecosystem, and implications for use of sciomyzids in biological control Author Williams, C. D. Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; Author Knutson, L. Author Gordh, G. Plant Protection & Quarantine, APHIS USDA, Raleigh, NC, USA; Author Mc Donnell, R. J. Department of Crop and Soil Science, Oregon State University, Corvallis, OR, USA; Author Kula, R. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Gates, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Buffington, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Broad, G. Department of Entomology, Natural History Museum, London, UK & Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; text Journal of Natural History 2025 2025-01-30 59 5 - 8 331 396 https://doi.org/10.1080/00222933.2024.2443125 journal article 10.1080/00222933.2024.2443125 1464-5262 14798887

Table 3. (Continued).

Mesoleptus n.sp. 5 Mesoleptus n.sp. 5 Mesoleptus sp. (misidentified	WRMM WRMM CFWM	Ditaeniella humilis Pherbellia obscura Atrichomelina	− − −	− − New York,	Bratt et al. (1969) Bratt et al. (1969) Foote et al. (1960)
as Atractodes sp.)		pubera		Michigan
“	JS	Elgiva cucularia	(P-1) 26.ix/6.x	S. Moravia	RozkoŠný (1967)
“	HKT	Tetanocera plebeja	(P-1) 7.iv/?	Idaho	Foote, unpub.
Atractodes sp. A	HKT	Pherbellia sp.	(P-1) 14.iv/?	New York	Bratt et al., 1964
Theroscopus sp. A	HKT	Dictya sp.	(P-5) 13.iii-30.iv/?	New York	Knutson and Neff, unpub.
“	HKT	Tetanocera	?	New York	Foote (1961)
		ferruginea
“	HKT	Tetanocera	(P-1) 5.iv/?	New York	Berg, unpub.
		fuscinervis
Theroscopus sp. A, B	HKT	Sepedon fuscipennis	(L,P) various	New York,	Neff and Berg (1966)
				Michigan
Theroscopus sp. B “	HKT HKT	Dictya sp. Tetanocera	13.iii-1.v −	New York New York	Knutson and Neff, unpub. Foote (1961)
		ferruginea
“	HKT	Tetanocera	(P-1) 30.iv/?	New York	Foote, unpub.
		fuscinervis
“ Theroscopus sp. C	HKT HKT	Sepedon sp. Tetanocera	(P-1) 18.ii/? −	New York New York	Berg, unpub. Foote (1961)
		ferruginea
Theroscopus sp. E	HKT	Dictya sp.	12.iv/?	New York	Berg, unpub.
Theroscopus sp. F	HKT	Sepedon armipes	(P-1) 6.xi/?	New York	Neff and Berg (1966)
Theroscopus sp. G Theroscopus sp. H	HKT HKT	Elgiva solicita Anticheta	31.v - 7.vi/? −	Belgium −	Knutson and Berg (1964) Knutson, unpub.
		melanosoma
Theroscopus sp. I	HKT	Dictya sp.	(P-2) 13.iii/?	New York	Neff, unpub.
Theroscopus sp. J	HKT	Pherbellia griseola	(P-1) 26.iv/?	New York	Bratt et al. (1969)
Theroscopus sp. K	HKT	Tetanocera plebeja	(L-?) 7.iv/?	Idaho	Foote, unpub.
Theroscopus sp. L	HKT	Tetanocera sp.	10.v/?	Idaho	Foote, unpub.
Theroscopus sp. M	HKT	Tetanocera sp.	10.v/?	Idaho	Foote, unpub.
Theroscopus sp. “	HKT HKT	Sepedon tenuicornis Tetanocera plebeja	10.ii/? (L-”few”)	New York −	Neff and Berg (1966) Trelka and Foote (1970)
Mesoleptus incessor	PL	Tetanocera arrogans	(P-1) “spring”	S. Moravia	RozkoŠný (1965)
Hemiteles sp.? (determination	JS	Elgiva cucularia	(P-1) 26.ix/6.x	S. Moravia	RozkoŠný (1967)
not certain)