First tabulation and analysis of natural enemies of snail-killing flies (Diptera: Sciomyzidae), their position in the fly / mollusc ecosystem, and implications for use of sciomyzids in biological control Author Williams, C. D. Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; Author Knutson, L. Author Gordh, G. Plant Protection & Quarantine, APHIS USDA, Raleigh, NC, USA; Author Mc Donnell, R. J. Department of Crop and Soil Science, Oregon State University, Corvallis, OR, USA; Author Kula, R. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Gates, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Buffington, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Broad, G. Department of Entomology, Natural History Museum, London, UK & Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; text Journal of Natural History 2025 2025-01-30 59 5 - 8 331 396 https://doi.org/10.1080/00222933.2024.2443125 journal article 10.1080/00222933.2024.2443125 1464-5262 14798887

Table 3. (Continued).

Tolmerus atricapillus	?	Adult	Pherbina coryleti	−	Wales	Clements and
						Skidmore 1998)
Empididae
Empis opaca	A	Adult (1)	Tetanocera silvatica	v	England	Hobby and Smith
Empis tessalata	A	Adult (1)	Limnia unguicornis	−	England	(1962) Hobby and Smith
“	A	Adult (1)	Renocera pallida	−	England	(1961) Hobby and Smith
						(1961)
Scathophagidae Unidentified species		Adult (several)	Pherbellia cinerella	−	Ireland	Gormally, (1987)
Tabanidae Hybomitra schineri 20	LLP	Larvae	Tetanocera ferruginea	−	Greece	Knutson (1965)
Orthoptera
Tettigoniidae
Conocephalus saltator		Adult?	Sepedonea spp.	27.v	Goias, Brazil	Mello, in litt.
Acari
Argiopidae
Tetragnatha sternalis		Adult	Protodictya chilensis	6.iv	Chile	Abercrombie (1970)
Araneae
Salticidae		Adult (No?)	Sepedon plumbella			Jackson et al. (1998)
Aves
Acarinidae
Araneus umbraticus		Adult	Pherbellia pallidiventris	3.viii	Norway	Greve and Ökland
						(1989)
Fringillidae
Leucosticte tephrocotis		Adult (1)	Pteromicra angustipennis	6.vi	Amchitka Is., Alaska	Higman, in litt.
Caprimulgidae Chordeiles minor	?	Adult (1)	Sciomyzidae, undet.	−	U.S.A.	Mosher, in litt.
Icteridae Euphagus carolinus		Adult (1)	Sciomyzidae, undet.	−	U.S.A.	Orians, in litt.
Pisces
Salmonidae
Salmo sp.		Larva (1)	Tetanocera sp.	v	nr. Fairbanks, Alaska	Loftus, in litt.
(C) Pathogens				Collected / Died
Fungi
Hypocreales Hirsutella citriformis	A	Adult	Sepedon aenescens 2	?	Philippines.	Rombach and
						Roberts (1989)
Entomophthora sp.		Adult (2)	Dictya pictipes	12.vii / 13.vii	New York	Valley and Berg
						(1977)
Nematoda