First tabulation and analysis of natural enemies of snail-killing flies (Diptera: Sciomyzidae), their position in the fly / mollusc ecosystem, and implications for use of sciomyzids in biological control Author Williams, C. D. Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; Author Knutson, L. Author Gordh, G. Plant Protection & Quarantine, APHIS USDA, Raleigh, NC, USA; Author Mc Donnell, R. J. Department of Crop and Soil Science, Oregon State University, Corvallis, OR, USA; Author Kula, R. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Gates, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Buffington, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Broad, G. Department of Entomology, Natural History Museum, London, UK & Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; text Journal of Natural History 2025 2025-01-30 59 5 - 8 331 396 https://doi.org/10.1080/00222933.2024.2443125 journal article 10.1080/00222933.2024.2443125 1464-5262 14798887

Table 3. (Continued).

Wingless proctotrupid	A	Tetanocera	−	Denmark	Lundbeck (1923)
		ferruginea
“		Sciomyza varia	(P-2) 10.ii.1955 / 3,10.iii	New York	Berg, unpub.
(B) Predators
Natural Enemy	Natural Enemy Identifier	Stage Attacked (Number) 19	Sciomyzidae	Dates Collected	Location	Reference
Hymenoptera
Sphecidae
Bembix amoena Bembix spinolae Ectemnius paucimaculatus “ “	A A A A A	Adult (1) Adult (1) Adult (1) Adult (4) Adult (1)	Tetanocera vicina Sepedon fuscipennis Dictya texensis Sepedon armipes Sepedon sp.	vii - viii − − − −	Wyoming Massachusetts Maryland Maryland Maryland	Evans (1966) Evans (1966) Krombein (1964) Krombein (1964) Krombein (1964)
Crabronidae
Dasyproctus bipunctatus	?	Adult (1)	Ethiolimnia lindneri	13.x - 7.xi	South Africa	Verbeke (1962)
jucundus
Crabro hilaris	A	Adult (1)	Dictya sp.			Matthews et al.
						(1979)
Crabo sp.		Adult (1)	Atrichomelina pubera	ix	Illinois	Specimen in USNM
Formicidae Pheidole megacephala	A	Larvae and Puparia	Sepedomerus macropus	−	Hawaii	Chock et al. (1961)
		(many)
Odonata Anisoptera naiads Zygoptera naiads	A A	Larvae (several) Larvae (several)	Sepedomerus macropus Sepedomerus macropus	− −	Hawaii Hawaii	Chock et al. (1961) Chock et al. (1961)
Coleoptera
Hydrophilidae Tropisternus sp.	A	Larvae (several, in	Sepedon fuscipennis	−	New York	Eckblad and Berg
		lab)				(1972)
Hemiptera
Veliidae Mesovelia mulsanti	A	1st instar larvae	Sepedomerus macropus	−	Hawaii	Chock et al. (1961)
		(several)
Notonectidae Notonecta sp.	A	3rd instar larva	Sepedon fuscipennis	−	New York	Eckblad and Berg
		(one)				(1972)
Diptera
Asilidae Andrenosoma atrum Dioctria lateralis Dioctria lateralis Dysmachus echinurus	A???	Adult Adult Adult Adult	Pherbellia obtusa Trypetoptera punctulata Coremacera marginata Pherbellia albocostata	− − − −	France Romania Romania USSR (Caucasus)	Musso (1970) Weinberg (1973) Weinberg (1973) Richter (1968)
Tolmerus atricapillus	A	Adult (1)	Dichetophora obliterata	5.ix	England	Hobby (1933)