First tabulation and analysis of natural enemies of snail-killing flies (Diptera: Sciomyzidae), their position in the fly / mollusc ecosystem, and implications for use of sciomyzids in biological control Author Williams, C. D. Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; Author Knutson, L. Author Gordh, G. Plant Protection & Quarantine, APHIS USDA, Raleigh, NC, USA; Author Mc Donnell, R. J. Department of Crop and Soil Science, Oregon State University, Corvallis, OR, USA; Author Kula, R. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Gates, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Buffington, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Broad, G. Department of Entomology, Natural History Museum, London, UK & Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; text Journal of Natural History 2025 2025-01-30 59 5 - 8 331 396 https://doi.org/10.1080/00222933.2024.2443125 journal article 10.1080/00222933.2024.2443125 1464-5262 14798887

Table 3. (Continued).

Phaenocarpa impugnata	JP	Anticheta	(2 L-3) 25.vi (4.vii)/ 25, 29. Denmark		Knutson (1966), J. Papp (1972)
		brevipennis	vii
“	JP	Anticheta	(4 L-2) 26.vii (7, 10.viii	Denmark	Knutson (1966), J. Papp (1972)
		brevipennis	stet)/30.viii, 4.ix
“	JP	Anticheta	(1 L) 2.ix (18.ix)/10.x	Denmark	Knutson (1966), J. Papp (1972)
		brevipennis
“	JP	Anticheta brevipennis 17	(1 L?) 2.ix.14.ix/ /3.vii.	Denmark	Knutson (1966), J. Papp (1972)
“	JP	Anticheta	3.v/3.vi	Denmark	Knutson (1966), J. Papp (1972)
		brevipennis
Unidentified		Colobaea americana	(P-7)?/18-25.v	New York	Knutson and Bratt, unpub.
Hymenoptera
Pteromalidae
Trichomalopsis?dubia	BDB	Pherbellia n. nana	(P-1)?/11.v	New York	Bratt et al. (1969) and Bratt, unpub.
Eupteromalus sp.	BDB	Colobaea americana	(2) 14.iv/27.iv, 14.v	New York	Knutson and Bratt, unpub.
“	BDB	Colobaea americana	(1) 28.iv/11.v	New York	Knutson and Bratt, unpub.
“	BDB	Pherbellia seticoxa	(P-1)?/28.iv	New York	Bratt et al. (1969) and Bratt, unpub.
“	BDB	Sepedon aenescens	(P)? [6-15]	Thailand	Beaver et al. (1977)
“	BDB	Sepedon ferruginosa	(P) x/? [6-15]	Thailand	Beaver et al. (1977)
“	BDB	Dichetophora sp.	?/5.viii	Australia	Berg, unpub.
Spalangia nigra	BDB	Atrichomelina	?/3.vi	New York	Foote et al. (1960)
		pubera
Spalangia sp.?		Atrichomelina			Universal Chalcidoidea Database
		pubera
Trichomalopsis sp.		Sepedon sp.			Universal Chalcidoidea Database
Trichomalopsis dubia		Pherbellia seticoxa			Universal Chalcidoidea Database
“ Trichomalopsis oryzae	A?	Dictya sp. Sepedon aenescens	−	Japan	Universal Chalcidoidea Database Yoneda (1986)
Urolepis rufipes	BDB	Sepedon armipes	?/20.xi	New York	Neff and Berg (1966)
“	A	Atrichomelina	13.vii.71	New York	Gibson, G. (2000)	USNM
		pubera
Hymenoptera
Chalcididae
‘ Chalcid’	A	Colobaea bifasciella		Denmark	Lundbeck (1923)
“	A	Colobaea punctata		Denmark	Lundbeck (1923)
Undetermined (Cephalobaris		Colobaea punctata	15.ii-20.iii/19.iii-26.v	Spain	Knutson and Bratt, unpub.
eskelundi Kryger?)
Order Hymenoptera
Unidentified Family Undetermined		Ilione albiseta 18		Ireland	Gormally, (1987)
“		Oidematops ferrugineus 18	?/14.ix	Ohio	Foote (1977)