First tabulation and analysis of natural enemies of snail-killing flies (Diptera: Sciomyzidae), their position in the fly / mollusc ecosystem, and implications for use of sciomyzids in biological control Author Williams, C. D. Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; Author Knutson, L. Author Gordh, G. Plant Protection & Quarantine, APHIS USDA, Raleigh, NC, USA; Author Mc Donnell, R. J. Department of Crop and Soil Science, Oregon State University, Corvallis, OR, USA; Author Kula, R. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Gates, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Buffington, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Broad, G. Department of Entomology, Natural History Museum, London, UK & Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; text Journal of Natural History 2025 2025-01-30 59 5 - 8 331 396 https://doi.org/10.1080/00222933.2024.2443125 journal article 10.1080/00222933.2024.2443125 1464-5262 14798887

Table 3. (Continued).

“ “	CFWM CFWM	Elgiva solicita Dictya sp. 12	?/vi, vii -	New York New York	O’Neill (1973) O’Neill (1973)
“	CFWM	Dictya sp.	29.iii/?	Florida	O’Neill (1973)
“ “	CFWM CFWM	Dictya sp. Tetanocera plebeja 13	?/vi, vii -	New York New York	O’Neill (1973) O’Neill (1973)
“	CFWM	Sciomyzidae (sp.)	iii, iv, xii/?	Florida, Louisiana	O’Neill (1973)
“	CFWM	Sciomyzidae (sp.)	summer to 29.x/?	Michigan,	O’Neill (1973)
				Minnesota
				Illinois, New York
Trichopria sp.	CFWM	Atrichomelina	(P-1) 19.viii/?	New York	Foote et al. (1960)
		pubera
Undetermined species	CFWM	Anticheta	(P-1) 3.v - 4.v	Denmark	Knutson (1966)
		brevipennis
3. LARVAL-PUPAL PARASITOIDS 14
Hymenoptera
Ichneumonidae
Mesoleptus declivus	CFWM	Atrichomelina	(P-24) 9.iv-21.viii/?	New York	Foote et al. (1960)
		pubera
‘‘	HKT	Dictya sp.	(P-4) 30.iv/?	New York	Knutson, unpub.
“	HKT	Ditaeniella humilis	(P-2) 20,25.vi/?	Idaho	Bratt et al. (1969) and Bratt, unpub.
“	HKT	Pherbellia albovaria	(P-several)/?	New York	Bratt et al. (1969) and Bratt, unpub.
“	HKT	Pherbellia griseola	(P-1) 7.iv/?	New York	Bratt et al. (1969) and Bratt, unpub.
“	HKT	Pherbellia n. nana	(P-1) 5.vii/?	New York	Bratt et al. (1969) and Bratt, unpub.
“	HKT	Pherbellia quadrata	(P-1) 21.iv/?	New York	Bratt et al. (1969) and Bratt, unpub.
“	HKT	Pherbellia s.	(P-6) 7-9.iv/?	New York	Bratt et al. (1969) and Bratt, unpub.
		maculata
“	HKT	Pteromicra	3.iv/?	New York	Foote, unpub.
		pectorosa
“	HKT	Sciomyza varia	(P-5) 27.vii / 21,22,23,27.	Michigan	Berg, unpub.
viii, 9.ix
“	HKT	Sciomyza varia	x/ 20-28 days later 23.xi/?	New York	Gower-Teece, unpub.
“	HKT	Sepedon armipes	3.iii/? “early spring”	New York	Neff and Berg (1966)	Same
“	HKT	Sepedon fuscipennis	(L,P) various dates	New York,	Neff and Berg (1966)
				Michigan
“		Sepedon fuscipennis	?/24.viii	New York	Ashmead (1901)	USNM
Mesoleptus sp. nr. declivus	WRMM	Anticheta	28.iv/6.v	New York	Knutson and Abercrombie (1977)
		melanosoma