First tabulation and analysis of natural enemies of snail-killing flies (Diptera: Sciomyzidae), their position in the fly / mollusc ecosystem, and implications for use of sciomyzids in biological control Author Williams, C. D. Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; Author Knutson, L. Author Gordh, G. Plant Protection & Quarantine, APHIS USDA, Raleigh, NC, USA; Author Mc Donnell, R. J. Department of Crop and Soil Science, Oregon State University, Corvallis, OR, USA; Author Kula, R. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Gates, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Buffington, M. Systematic Entomology Laboratory, USDA-ARS, Smithsonian Institution, Washington, DC, USA; Author Broad, G. Department of Entomology, Natural History Museum, London, UK & Biological and Environmental Sciences, Liverpool John Moores University, Liverpool, UK; text Journal of Natural History 2025 2025-01-30 59 5 - 8 331 396 https://doi.org/10.1080/00222933.2024.2443125 journal article 10.1080/00222933.2024.2443125 1464-5262 14798887

Table 3. (Continued).

“			Sepedon sphegea			Universal Chalcidoidea Database
Trichogrammatoidea bactrae “ Trichogrammatoidea nana	??		Tetanocera sp. Sepedon aenescens 1 Sepedon aenescens 1 Sepedon aenescens 1	??	? India	Universal Chalcidoidea Database Hayat and Subba Rao (1986) Nagarkatti and Nagaraja (1977) Universal Chalcidoidea Database
Trichogrammatoidea
simmondsi Unidentified “	? A A		Sepedon ‘angularis’ Sciomyza aristalis 8 Sepedon spinipes 9	??	New York Denmark	Universal Chalcidoidea Database Foote (1959) Neff and Berg (1966)
2. PUPAL PARASITOIDS
Hymenoptera
Diapriidae
Spilomicrus barnesi	JWE and	DSH	Neolimnia tranquilla	(P-35) 18.iii, 4.iv, 26.vii,	New Zealand	Barnes (1979) and Early and Horning
						(1978)
				7xii/ [333]
Spilomicrus sp.	CFWM		Sepedonea trichotypa 10	(L3-1) 16.iv(19.iv)/19.v [4]	Argentina	Abercrombie (1970)
“ Trichopria atrichomelinae	CFWM CFWM		Sepedonea trichotypa 10 Atrichomelina pubera 11	(P-1) 16.iv/16.v [7] 20-27.vii/25-26.viii 9,11,11	Argentina New York	Abercrombie (1970) Muesebeck, (1972) and O’Neill (1973)
“	CFWM?		Sepedon fuscipennis 12	Oviposition, no emergence	New York	O’Neill (1973)
“ “	CFWM? CFWM?		Elgiva solicita 12 Dictya sp. 12	“ “	New York New York	O’Neill (1973) O’Neill (1973)
Trichopria popei	?		Dictya floridensis	(P-4) 29.iii/? [19]	Florida	Valley and Berg (1977)
“	CFWM		Dictya sp.	(P-1) 13.viii/31.viii [5]	Minnesota	Knutson and Berg (1963)
“	CFWM		Dictya sp.	(P-1) 29.x/1.xi [10]	New York	Knutson and Berg (1963)
“	CFWM		Elgiva solicita	(P-1) 2.viii/12.viii [16]	New York	Knutson and Berg (1963)
“	CFWM		Sepedon fuscipennis	(P-1) 28.vi/1.vii [30]	Michigan	Muesebeck (1949) and Berg (1953) and	CU
						Knutson and Berg 1963)
“	CFWM		Sepedon fuscipennis	(P-1) 16.vii/21.vii [17]	Michigan	Berg (1953) and Knutson and Berg 1963)	CU
“	CFWM		Sepedon fuscipennis	?/vi, vii	New York	O’Neill (1973)
“	CFWM		Atrichomelina pubera 12	-	New York	O’Neill (1973)
“	CFWM		Sepedon fuscipennis 12	-	New York	O’Neill (1973)
“	CFWM		Elgiva solicita 12	-	New York	O’Neill (1973)