First record of Metapolystoma (Monogenea: Polystomatidae) from Boophis tree frogs in Madagascar, with the description of five new species Author Landman, Willem Unit for Environmental Sciences and Management, North-West University, Potchefstroom Campus, Private Bag X 6001, Potchefstroom, 2520, South Africa Willie.Landman@nwu.ac.za Author Verneau, Olivier Unit for Environmental Sciences and Management, North-West University, Potchefstroom Campus, Private Bag X 6001, Potchefstroom, 2520, South Africa & University of Perpignan Via Domitia Centre de Formation et de Recherche sur les Environnements M´editerran´eens, UMR 5110, F- 66860, Perpignan, France & CNRS, Centre de Formation et de Recherche sur les Environnements M´editerran´eens, UMR 5110, F- 66860, Perpignan, France Author Raharivololoniaina, Liliane Department of Animal Biology, University of Antananarivo, BP 566, Antananarivo 101, Madagascar Author Preez, Louis du Unit for Environmental Sciences and Management, North-West University, Potchefstroom Campus, Private Bag X 6001, Potchefstroom, 2520, South Africa & South African Institute for Aquatic Biodiversity, Private Bag 1015, Makhanda, 6140, South Africa text International Journal for Parasitology: Parasites and Wildlife 2021 2021-04-30 14 161 178 http://dx.doi.org/10.1016/j.ijppaw.2021.01.012 journal article 292465 10.1016/j.ijppaw.2021.01.012 3afcda9a-df8e-47e9-9cab-f8ac8fe18e5f 2213-2244 PMC8056147 33898217 10668146

Table 4 Mean genetic distances (below diagonal) and total character differences (above diagonal) between species groups as inferred from comparisons of 18S rDNA sequences (1931 characters).

		1	2	3	4	5	6	7	8	9	10	11
1	M. brygoonis		1	2	2	2	11	10	13	10	9	25
2	M. falcatum n. sp. (B. d.)	0,001		1	1	1	10	9	12	9	8	24
3	M. multiova n. sp. (B. o.)	0,001	0001		0	2	11	10	13	10	9	25
4	M. theroni n. sp. (B. m)	0,001	0001	0,000		2	11	10	13	10	9	25
5	M. cachani	0,001	0001	0,001	0001		9	8	11	10	9	23
6	P. occipitalis	0,006	0005	0,006	0006	0,005		6	16	15	14	28
7	P. dawiekoki	0,005	0005	0,005	0005	0,004	0003		16	15	14	28
8	P. claudecombesi	0,007	0006	0,007	0007	0,006	0008	0,008		11	10	24
9	P. marmorati	0,005	0005	0,005	0005	0,005	0008	0,008	0006		1	25
10	P. testimagna	0,005	0004	0,005	0005	0,005	0007	0,007	0005	0,001		24
11	P. integerrimum	0,013	0013	0,013	0013	0,012	0015	0,015	0013	0,013	0,0126